Astragalus atratus

S. Watson

Botany (Fortieth Parallel), 69, plate 11. 1871.

Common names: Mourning milkvetch
Endemic
Treatment appears in FNA Volume 11.

Plants caulescent, (1–)3–24(–30) cm, strigulose. Stems prostrate to ascending, strigulose. Leaves (1.5–)3–14(–14.5) cm; stipules distinct, 1–5 mm, papery-scarious at proximal nodes, subherbaceous at distal nodes; leaflets 7–15, blades oblong-oblanceolate, elliptic, oval-oblong, linear-oblong, linear-elliptic, or linear-setaceous, 1.5–16 mm, apex obtuse, retuse, subacute, or emar­ginate, surfaces strigulose abaxially, strigulose or glabrous adaxially; terminal leaflet sometimes decurrent distally, jointed to or continuous with rachis. Peduncles incurved-ascending, (1.5–)3–16.5 cm. Racemes (2–)5–18-flowered, flowers nodding; axis (1.5–)3–17 cm in fruit; bracts (0.8–)1–2.5 mm; bracteoles usually 0. Pedicels (1.5–)2–6.5 mm. Flowers 6.3–13.4 mm; calyx 3–7.6 mm, strigulose, tube 2.3–5.6 mm, lobes subulate, 0.7–2.6 mm; corolla whitish or dirty white, sometimes veined, tipped, margined, or suffused with purple, or faintly tinged lavender; keel 6–10 mm. Legumes pendu­lous, green, purplish, or mottled, decurved or straight, linear-oblong, linear-oblanceoloid, or narrowly ellipsoid, compressed, ventral suture convex in profile or at least beak declined from body, 10–22 × 2.3–4.3 mm, unilocular or bilocular, thinly fleshy becoming papery or leathery, strigulose; septum (0 or)0.6–2.7 mm wide; stipe 0–2 mm, concealed within calyx when present. Seeds 10–29.

Distribution

w United States.

Discussion

Varieties 4 (4 in the flora).

R. C. Barneby (1964) related that the main pattern underlying the complex racial differentiation in Astragalus atratus is one of northern and southern branches relating to the Snake-Humboldt divide in north­eastern Nevada. Populations there and northward have subentire wing petals and unilocular fruits. South­ward, the wings are toothed or lobed, and the fruit is nearly bilocular.

Selected References

None.

Key

1 Legumes laterally compressed, bilocular, stipes 0–0.7 mm; Death Valley, Inyo County, California. Astragalus atratus var. mensanus
1 Legumes ± dorsiventrally compressed, unilocu­lar or bilocular, stipes 0.3–2 mm; c Nevada to e Oregon and sw Idaho. > 2
2 Legumes sub-bilocular, leathery, septae 0.6–1.3 mm wide; terminal leaflets jointed to rachises; c, nc Nevada. Astragalus atratus var. atratus
2 Legumes unilocular, leathery or papery, septae obsolete or to 0.2 mm wide; terminal leaflets jointed to or continuous with rachises; Idaho, Oregon, extreme ne Nevada. > 3
3 Leaflet blades narrowly oblong-oblanceolate to linear-oblong or oval, 3–13 mm, terminal leaflets jointed to rachises; legumes leathery; n side of Snake River Plains in Blaine, Camas, and Lincoln counties, Idaho. Astragalus atratus var. inseptus
3 Leaflet blades linear-elliptic, 1.5–7(–10) mm, terminal leaflets continuous with rachises or reduced to slight dilation of rachis tips; legumes papery; e Oregon, extreme sw Idaho (Owyhee to Washington counties), ne Nevada. Astragalus atratus var. owyheensis