Rhinanthus

Linnaeus

Sp. Pl. 2: 603. 1753.

Common names: Yellow rattle
Etymology: Greek rhis, nose or snout, and anthos, flower, alluding to protruding adaxial lip of corolla
Treatment appears in FNA Volume 17. Treatment on page 504. Mentioned on page 456, 458.

Herbs, annual; hemiparasitic. Stems erect, not fleshy, glabrous, hairy on 2 opposite sides. Leaves cauline, opposite; petiole present or absent; blade not fleshy to +/- fleshy, not leathery, margins coarsely toothed distally. Inflorescences terminal, racemes, +/- secund; bracts present. Pedicels present; bracteoles absent. Flowers: sepals 4, calyx bilaterally symmetric, flattened laterally, ovate to suborbiculate, accrescent in fruit, lobes deltate; petals 5, corolla yellow or yellowish [bronze to bluish], compressed, strongly bilabiate, narrowly campanulate, abaxial lobes 3, adaxial 2, adaxial lip galeate, ovate, obtuse, entire, with subapical tooth on each side; stamens 4, didynamous, filaments lanate; staminode 0; ovary 2-locular, placentation axile; stigma capitate. Capsules: dehiscence loculicidal longitudinally. Seeds 2–10[+], brown, kidney-shaped, flattened, wings present, absent in some R. major. x = 11.

Distribution

North America, Europe, n Asia.

Discussion

Species ca. 50 (2 in the flora).

Although the taxonomy of Rhinanthus in the Old World is complex, the taxa found in North America are reasonably distinct. The subapical teeth on the adaxial corolla lip have been described as galea or nipples; the term teeth is used in this account. Leaf measurements are for mid-stem leaves; calyx features are for post-anthesis calyces.

The European species Rhinanthus serotinus (Schönheit ex Halácsy & H. Braun) Oborný has been reported from Maine (E. Hultén and M. Fries 1986); the authors have found no specimens to substantiate this report. If it were to be encountered in North America, it would key to R. major, from which it can be distinguished by having narrowly triangular bracts and glabrous calyces as in R. minor.

Selected References

None.

Key

1 Corollas 15–20 mm; teeth of galea of corolla 1.5–2.5 mm; calyces villous, glabrescent. Rhinanthus major
1 Corollas 13–15 mm; teeth of galea of corolla (0.4–)0.6–1 mm; calyces glabrous, scabrid, or puberulent. Rhinanthus minor