Echinochloa colona
Plants annual; erect or decumbent, cespitose or spreading, rooting from the lower cauline nodes. Culms 10-70 cm; lower nodes glabrous or hispid, hairs appressed; upper nodes glabrous. Sheaths glabrous; ligules absent, ligule region frequently brown-purple; blades 8-22 cm long, 3-6(10) mm wide, mostly glabrous, sometimes hispid, hairs papillose-based on or near the margins. Panicles 2-12 cm, erect, rachises glabrous or sparsely hispid; primary branches 5-10, 0.7-2(4) cm, erect to ascending, spikelike, somewhat distant, without secondary branches, axes glabrous or sparsely hispid, hairs 1.5-2.5 mm, papillose-based. Spikelets 2-3 mm, disarticulating at maturity, pubescent to hispid, hairs usually not papillose-based, tips acute to cuspidate. Lower glumes about 1/2 as long as the spikelets; upper glumes about as long as the spikelets; lower florets usually sterile, occasionally staminate; lower lemmas unawned, similar to the upper glumes; lower paleas subequal to the lemmas; upper lemmas 2.6-2.9 mm, not or scarcely exceeding the upper glumes, elliptic, coriaceous portion rounded distally, passing abruptly into a sharply differentiated, membranous, soon-withering tip; anthers 0.7-0.8 mm. Caryopses 1.2-1.6 mm, whitish; embryos 63-83% as long as the caryopses. 2n = 54.
Distribution
Mont., Oreg., Wash., Pacific Islands (Hawaii), Fla., N.J., N.Mex., Tex., La., Tenn., N.C., S.C., Pa., Va., Calif., Puerto Rico, Virgin Islands, Ala., Ark., Vt., Ill., Ga., Mass., Ariz., Md., Okla., Mo., Kans., Miss., Ky.
Discussion
Echinochloa colona is widespread in tropical and subtropical regions. It is adventive and weedy in North America, growing in low-lying, damp to wet, disturbed areas, including rice fields. The unbranched, rather widely-spaced panicle branches make this one of the easier species of Echinochloa to recognize.
Hitchcock (1913) considered that 'colonum' was a non-declining contraction, but dictionaries of Linnaeus' time treated it as a declining adjective. Because Linnaeus was the first to name the species (as "Panicum colonum"), it seems best to follow the practice considered correct in his day; hence "E. colona".
Selected References
None.