Diphasiastrum complanatum

(Linnaeus) Holub

Preslia 47: 108. 1975.

Common names: Northern running-pine lycopode aplati
Basionym: Lycopodium complanatum Linnaeus Sp. Pl. 2: 1104. 1753
Synonyms: Lycopodium complanatum var. canadense Victorin
Treatment appears in FNA Volume 2.

Horizontal stems on substrate surface or shallowly buried in litter, 1.1–2.2 mm wide; leaves appressed, linear to narrowly lanceolate, 1.4–4 × 0.5–1.2 mm, apices acute. Upright shoots 8–44 cm, branching irregularly successively to 5 times; leaves on upright main stem appressed with decurrent base, narrowly lanceolate, 1.2–3.2 × 0.5–1.1 mm, apex acute to acuminate. Branchlets flat in cross section, narrowly bladelike, 1.8–4 mm wide, annual bud constrictions abrupt and conspicuous; upperside green, faintly shiny, flat; underside dull, pale, flat. Leaves on branchlets 4-ranked; upperside leaves appressed, linear-lanceolate, free portion of blades 0.7–2 × 0.5–1.2 mm; lateral leaves appressed, 2.6–7.3 × 0.8–2.1 mm; underside leaves weakly developed, appressed, narrowly deltate, 0.7–1.5 × 0.4–0.9 mm. Peduncles 1–2 on each upright shoot, 0.5–8.5 × 0.4–0.9 cm; leaves spirally arranged to nearly whorled, linear-lanceolate, 1.4–4.1 × 0.4–1 mm, apex acute to blunt. Stalks forked at uniform distances. Strobili 1–2(–4), 8.3–32 × 2–3 mm, apex blunt, sterile tip absent. Sporophylls broadly deltate to nearly cordate, 2–3 × 2–2.4 mm, apex abruptly tapering. 2n = 46.

Habitat: Dry open coniferous or mixed forest alpine slopes
Elevation: 0–2000 m


V2 574-distribution-map.gif

Greenland, St. Pierre and Miquelon, Alta., B.C., Man., N.B., Nfld. and Labr. (Nfld.), N.W.T., N.S., Nunavut, Ont., P.E.I., Que., Sask., Yukon, Alaska, Idaho, Maine, Mich., Minn., Mont., N.H., N.Y., Vt., Wash., Wis., Wyo., circumboreal.


Diphasiastrum complanatum forms a hybrid with D. digitatum that is seemingly uncommon and has never received a binomial designation. It is probably far more common than collections indicate, however. Superficially, the hybrid resembles both parents and is often confused with them. Collections are known from Ontario, Quebec, Connecticut, Maine, Michigan, Minnesota, New Hampshire, Vermont, and Wisconsin.

Selected References


Lower Taxa

... more about "Diphasiastrum complanatum"
Warren H. Wagner Jr. +  and Joseph M. Beitel +
(Linnaeus) Holub +
Lycopodium complanatum +
Northern running-pine +  and lycopode aplati +
Greenland +, St. Pierre and Miquelon +, Alta. +, B.C. +, Man. +, N.B. +, Nfld. and Labr. (Nfld.) +, N.W.T. +, N.S. +, Nunavut +, Ont. +, P.E.I. +, Que. +, Sask. +, Yukon +, Alaska +, Idaho +, Maine +, Mich. +, Minn. +, Mont. +, N.H. +, N.Y. +, Vt. +, Wash. +, Wis. +, Wyo. +  and circumboreal. +
0–2000 m +
Dry open coniferous or mixed forest alpine slopes +
Illustrated +
Lycopodium complanatum var. canadense +
Diphasiastrum complanatum +
Diphasiastrum +
species +