Difference between revisions of "Woodsia cochisensis"
Contr. Univ. Michigan Herb. 19: 54. 1993.
Common names: Cochise cliff fern
Treatment appears in FNA Volume 2.
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| − | --><span class="statement" id="st- | + | --><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact, erect to ascending, with a few persistent petiole bases of unequal lengths; scales often uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. <b>Leaves</b> 5–25 × 1.5–6 cm. <b>Petiole</b> light brown or straw-colored throughout when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered. <b>Blade</b> narrowly lanceolate to ovate, pinnate-pinnatifid to 2-pinnate proximally, sparsely to moderately glandular, never viscid; glandular hairs with thin stalks and slightly expanded tips; rachis with glandular hairs and occasional hairlike scales. <b>Pinnae</b> ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex, occasionally attenuate; largest pinnae with 4–9 pairs of pinnules; abaxial and adaxial surfaces glandular, lacking nonglandular hairs or scales. <b>Pinnules</b> dentate, often shallowly lobed; margins lustrous adaxially, usually thickened, lacking cilia but sparsely glandular, with occasional 1–2-celled translucent projections. <b>Vein</b> tips enlarged to form whitish hydathodes visible adaxially. <b>Indusia</b> of relatively broad segments; segments multiseriate most of length, usually divided and uniseriate distally, composed of ± isodiametric cells, often surpassing mature sporangia. <b>Spores</b> averaging 43–49 µm. <b>2n</b> = 152.</span><!-- |
-->{{Treatment/Body | -->{{Treatment/Body | ||
| + | |phenology=Sporulating late spring–fall. | ||
|habitat=Shaded ledges and alcoves near springs and seeps, usually on granitic or volcanic substrates | |habitat=Shaded ledges and alcoves near springs and seeps, usually on granitic or volcanic substrates | ||
|elevation=1000–2200 m | |elevation=1000–2200 m | ||
|distribution=Ariz.;N.Mex.;n Mexico. | |distribution=Ariz.;N.Mex.;n Mexico. | ||
| − | |discussion=<p>Woodsia cochisensis traditionally has been identified as W. plummerae or (rarely) W. mexicana Fée. It is readily separated from W. plummerae by the characteristics given in the key, and from North American members of the mexicana group (W. phillipsii and W. neomexicana) by having indusial segments that are broad and nonfilamentous at the base. Woodsia cochisensis is less glandular than typical W. mexicana from northeastern Mexico and is further distinguished from that species by the thickened, lustrous pinnule margins and well-developed hydathodes. Isozyme and chromosome studies suggest that W. cochisensis is an allotetraploid that may have originated through hybridization between W. phillipsii and an undescribed Mexican diploid (M. D. Windham 1993). It crosses with the former species to produce sterile triploids of intermediate morphology.</p> | + | |discussion=<p><i>Woodsia cochisensis</i> traditionally has been identified as <i>W. plummerae</i> or (rarely) W. mexicana Fée. It is readily separated from <i>W. plummerae</i> by the characteristics given in the key, and from North American members of the mexicana group (<i>W. phillipsii</i> and <i>W. neomexicana</i>) by having indusial segments that are broad and nonfilamentous at the base. <i>Woodsia cochisensis</i> is less glandular than typical W. mexicana from northeastern Mexico and is further distinguished from that species by the thickened, lustrous pinnule margins and well-developed hydathodes. Isozyme and chromosome studies suggest that <i>W. cochisensis</i> is an allotetraploid that may have originated through hybridization between <i>W. phillipsii</i> and an undescribed Mexican diploid (M. D. Windham 1993). It crosses with the former species to produce sterile triploids of intermediate morphology.</p> |
|tables= | |tables= | ||
|references= | |references= | ||
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-->{{#Taxon: | -->{{#Taxon: | ||
name=Woodsia cochisensis | name=Woodsia cochisensis | ||
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|authority=Windham | |authority=Windham | ||
|rank=species | |rank=species | ||
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|basionyms= | |basionyms= | ||
|family=Dryopteridaceae | |family=Dryopteridaceae | ||
| + | |phenology=Sporulating late spring–fall. | ||
|habitat=Shaded ledges and alcoves near springs and seeps, usually on granitic or volcanic substrates | |habitat=Shaded ledges and alcoves near springs and seeps, usually on granitic or volcanic substrates | ||
|elevation=1000–2200 m | |elevation=1000–2200 m | ||
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|publication year=1993 | |publication year=1993 | ||
|special status= | |special status= | ||
| − | |source xml=https:// | + | |source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_370.xml |
|genus=Woodsia | |genus=Woodsia | ||
|species=Woodsia cochisensis | |species=Woodsia cochisensis | ||
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-->[[Category:Treatment]][[Category:Woodsia]] | -->[[Category:Treatment]][[Category:Woodsia]] | ||
Latest revision as of 21:23, 5 November 2020
Stems compact, erect to ascending, with a few persistent petiole bases of unequal lengths; scales often uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. Leaves 5–25 × 1.5–6 cm. Petiole light brown or straw-colored throughout when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered. Blade narrowly lanceolate to ovate, pinnate-pinnatifid to 2-pinnate proximally, sparsely to moderately glandular, never viscid; glandular hairs with thin stalks and slightly expanded tips; rachis with glandular hairs and occasional hairlike scales. Pinnae ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex, occasionally attenuate; largest pinnae with 4–9 pairs of pinnules; abaxial and adaxial surfaces glandular, lacking nonglandular hairs or scales. Pinnules dentate, often shallowly lobed; margins lustrous adaxially, usually thickened, lacking cilia but sparsely glandular, with occasional 1–2-celled translucent projections. Vein tips enlarged to form whitish hydathodes visible adaxially. Indusia of relatively broad segments; segments multiseriate most of length, usually divided and uniseriate distally, composed of ± isodiametric cells, often surpassing mature sporangia. Spores averaging 43–49 µm. 2n = 152.
Phenology: Sporulating late spring–fall.
Habitat: Shaded ledges and alcoves near springs and seeps, usually on granitic or volcanic substrates
Elevation: 1000–2200 m
Distribution
Ariz., N.Mex., n Mexico.
Discussion
Woodsia cochisensis traditionally has been identified as W. plummerae or (rarely) W. mexicana Fée. It is readily separated from W. plummerae by the characteristics given in the key, and from North American members of the mexicana group (W. phillipsii and W. neomexicana) by having indusial segments that are broad and nonfilamentous at the base. Woodsia cochisensis is less glandular than typical W. mexicana from northeastern Mexico and is further distinguished from that species by the thickened, lustrous pinnule margins and well-developed hydathodes. Isozyme and chromosome studies suggest that W. cochisensis is an allotetraploid that may have originated through hybridization between W. phillipsii and an undescribed Mexican diploid (M. D. Windham 1993). It crosses with the former species to produce sterile triploids of intermediate morphology.
Selected References
None.
Lower Taxa
None.