Difference between revisions of "Helleborus"

Linnaeus

Sp. Pl. 1: 557. 175.

,

Gen. Pl. ed. 5, 244. 1754.

Common names: Hellebore hellébore
Etymology: Greek, helleborus, ancient name for this plant
Treatment appears in FNA Volume 3.
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|title=Sp. Pl.
 
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|title=Gen. Pl. ed.
 
|title=Gen. Pl. ed.
 
|place=5, 244. 1754
 
|place=5, 244. 1754
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--><span class="statement" id="st-d0_s0" data-properties="herb duration;rhizome fragility;rhizome height or length or size"><b>Herbs </b>[subshrubs], perennial, from tough, short rhizomes [rhizomes absent].</span> <span class="statement" id="st-d0_s1" data-properties="leaf position;leaf position;basal leaf size;basal leaf architecture"><b>Leaves </b>basal and cauline, basal leaf much larger [all leaves cauline], petiolate;</span> <span class="statement" id="st-d0_s2" data-properties="cauline leaf arrangement">cauline leaves alternate.</span> <span class="statement" id="st-d0_s3" data-properties="leaf-blade architecture;leaf-blade shape;lobe shape;lobe shape;lobe shape;lobe shape;margin shape"><b>Leaf-</b>blade pedately or palmately compound or deeply parted [undivided], lobes narrowly elliptic to oblanceolate or lanceolate, margins sharply toothed [entire].</span> <span class="statement" id="st-d0_s4" data-properties="inflorescence position or structure subtype;cyme architecture;cyme some measurement;flower arrangement;flower arrangement"><b>Inflorescences </b>terminal, 3-4-flowered cymes, to 25 cm or flowers solitary or paired;</span> <span class="statement" id="st-d0_s5" data-properties="beak architecture or shape;beak shape">bracts ±leaflike, divided, not forming involucre.</span> <span class="statement" id="st-d0_s6" data-properties="flower reproduction;flower architecture or shape"><b>Flowers </b>bisexual, radially symmetric;</span> <span class="statement" id="st-d0_s7" data-properties="sepal duration;sepal coloration;sepal shape;sepal shape;sepal shape;sepal shape;sepal atypical some measurement;sepal some measurement;fruit quantity">sepals persistent in fruit [not persistent], 5, yellowish green [white, pink, or purple], plane, ovate to elliptic, 19-30 (-50) mm;</span> <span class="statement" id="st-d0_s8" data-properties="petal quantity;petal fusion;petal coloration;petal coloration;petal shape;petal architecture;petal shape;petal some measurement">petals 5-15, distinct, green or brown, funnel-shaped, ± 2-lipped, clawed, 4-8 mm;</span> <span class="statement" id="st-d0_s9" data-properties="">nectary in center of funnel;</span> <span class="statement" id="st-d0_s10" data-properties="stamen quantity">stamens 30-60;</span> <span class="statement" id="st-d0_s11" data-properties="filament shape">filaments filiform;</span> <span class="statement" id="st-d0_s12" data-properties="staminode presence">staminodes absent between stamens and pistils;</span> <span class="statement" id="st-d0_s13" data-properties="pistil atypical quantity;pistil atypical quantity;pistil quantity;pistil architecture;pistil fusion">pistils [2-] 3-6 [-10], simple, proximally connate [distinct or completely connate];</span> <span class="statement" id="st-d0_s14" data-properties="ovule quantity">ovules several per pistil;</span> <span class="statement" id="st-d0_s15" data-properties="style presence">style present.</span> <span class="statement" id="st-d0_s16" data-properties="fruits follicle architecture;fruits follicle architecture;fruits follicle shape;vein prominence;vein dehiscence or orientation"><b>Fruits </b>follicles [capsules], aggregate, sessile, oblong, sides with prominent transverse veins;</span> <span class="statement" id="st-d0_s17" data-properties="beak position or structure subtype;beak course;beak some measurement">beak terminal, straight, 5-15 mm.</span> <span class="statement" id="st-d0_s18" data-properties=""><b>Seeds </b>usually ± carinate.</span> <span class="statement" id="st-d0_s19" data-properties="seed shape;x chromosome quantity">x = 8.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Herbs </b>[subshrubs], perennial, from tough, short rhizomes [rhizomes absent]. <b>Leaves</b> basal and cauline, basal leaf much larger [all leaves cauline], petiolate; cauline leaves alternate. <b>Leaf</b> blade pedately or palmately compound or deeply parted [undivided], lobes narrowly elliptic to oblanceolate or lanceolate, margins sharply toothed [entire]. <b>Inflorescences</b> terminal, 3-4-flowered cymes, to 25 cm or flowers solitary or paired; bracts ±leaflike, divided, not forming involucre. <b>Flowers</b> bisexual, radially symmetric; sepals persistent in fruit [not persistent], 5, yellowish green [white, pink, or purple], plane, ovate to elliptic, 19-30(-50) mm; petals 5-15, distinct, green or brown, funnel-shaped, ± 2-lipped, clawed, 4-8 mm; nectary in center of "funnel"; stamens 30-60; filaments filiform; staminodes absent between stamens and pistils; pistils [2-]3-6[-10], simple, proximally connate [distinct or completely connate]; ovules several per pistil; style present. <b>Fruits</b> follicles [capsules], aggregate, sessile, oblong, sides with prominent transverse veins; beak terminal, straight, 5-15 mm. <b>Seeds</b> usually ± carinate. <b>x</b> = 8.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
|distribution=North America;Europe;Asia (in Asia Minor and Tibet)
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|distribution=North America;Europe;Asia (in Asia Minor and Tibet).
 
|discussion=<p>Species ca. 25 (1 in the flora).</p><!--
 
|discussion=<p>Species ca. 25 (1 in the flora).</p><!--
--><p>Although other species of Helleborus are grown as ornamentals, only the green-flowered H. viridis appears to persist after cultivation. Helleborus niger Linnaeus (Christmas-rose) is a more popular ornamental because of its showy, white to pinkish flowers. It does not appear to persist away from cultivation; it was reported as an escape in 1880 at Sennet, New York, and in 1919 in Washtenaw County, Michigan (R. S. Mitchell and J. K. Dean 1982; E. G. Voss 1972+, vol. 2). Helleborus niger can be distinguished from H. viridis by its flower color and its simple, distal cauline leaves with entire margins.</p><!--
+
--><p>Although other species of <i>Helleborus</i> are grown as ornamentals, only the green-flowered <i>H. viridis</i> appears to persist after cultivation. <i>Helleborus</i> niger Linnaeus (Christmas-rose) is a more popular ornamental because of its showy, white to pinkish flowers. It does not appear to persist away from cultivation; it was reported as an escape in 1880 at Sennet, New York, and in 1919 in Washtenaw County, Michigan (R. S. Mitchell and J. K. Dean 1982; E. G. Voss 1972+, vol. 2). <i>Helleborus</i> niger can be distinguished from <i>H. viridis</i> by its flower color and its simple, distal cauline leaves with entire margins.</p><!--
--><p>Both living and dried plants of all species of Helleborus are extremely poisonous. Plants contain a cardiac glycoside (helleborin), which acts directly on the heart muscle, causing convulsions, delirium, and sometimes death. Poisoning from contaminated hay has been known to cause livestock fatalities in some areas (R. S. Mitchell and J. K. Dean 1982).</p>
+
--><p>Both living and dried plants of all species of <i>Helleborus</i> are extremely poisonous. Plants contain a cardiac glycoside (helleborin), which acts directly on the heart muscle, causing convulsions, delirium, and sometimes death. Poisoning from contaminated hay has been known to cause livestock fatalities in some areas (R. S. Mitchell and J. K. Dean 1982).</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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|basionyms=
 
|basionyms=
 
|family=Ranunculaceae
 
|family=Ranunculaceae
|distribution=North America;Europe;Asia (in Asia Minor and Tibet)
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|distribution=North America;Europe;Asia (in Asia Minor and Tibet).
 
|reference=None
 
|reference=None
 
|publication title=Sp. Pl.;Gen. Pl. ed.
 
|publication title=Sp. Pl.;Gen. Pl. ed.
 
|publication year=;1754
 
|publication year=;1754
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V3/V3_192.xml
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|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V3/V3_192.xml
 
|genus=Helleborus
 
|genus=Helleborus
|basal leaf architecture=petiolate
 
|basal leaf size=larger
 
|beak architecture or shape=leaflike
 
|beak course=straight
 
|beak position or structure subtype=terminal
 
|beak shape=divided
 
|beak some measurement=5mm;15mm
 
|cauline leaf arrangement=alternate
 
|cyme architecture=3-4-flowered
 
|cyme some measurement=0cm;25cm
 
|filament shape=filiform
 
|flower architecture or shape=symmetric
 
|flower arrangement=paired;solitary
 
|flower reproduction=bisexual
 
|fruit quantity=5
 
|fruits follicle architecture=sessile;aggregate
 
|fruits follicle shape=oblong
 
|herb duration=perennial
 
|inflorescence position or structure subtype=terminal
 
|leaf position=cauline;basal
 
|leaf-blade architecture=compound
 
|leaf-blade shape=parted
 
|lobe shape=narrowly elliptic;oblanceolate or lanceolate
 
|margin shape=toothed
 
|ovule quantity=several
 
|petal architecture=2-lipped
 
|petal coloration=brown;green
 
|petal fusion=distinct
 
|petal quantity=5;15
 
|petal shape=clawed;funnel--shaped
 
|petal some measurement=4mm;8mm
 
|pistil architecture=simple
 
|pistil atypical quantity=6;10
 
|pistil fusion=connate
 
|pistil quantity=3;6
 
|rhizome fragility=tough
 
|rhizome height or length or size=short
 
|seed shape=carinate
 
|sepal atypical some measurement=30mm;50mm
 
|sepal coloration=yellowish green
 
|sepal duration=persistent
 
|sepal shape=ovate;elliptic
 
|sepal some measurement=19mm;30mm
 
|stamen quantity=30;60
 
|staminode presence=absent
 
|style presence=absent
 
|vein dehiscence or orientation=transverse
 
|vein prominence=prominent
 
|x chromosome quantity=8
 
 
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-->[[Category:Treatment]][[Category:Ranunculaceae]]
 
-->[[Category:Treatment]][[Category:Ranunculaceae]]

Latest revision as of 22:46, 5 November 2020

Herbs [subshrubs], perennial, from tough, short rhizomes [rhizomes absent]. Leaves basal and cauline, basal leaf much larger [all leaves cauline], petiolate; cauline leaves alternate. Leaf blade pedately or palmately compound or deeply parted [undivided], lobes narrowly elliptic to oblanceolate or lanceolate, margins sharply toothed [entire]. Inflorescences terminal, 3-4-flowered cymes, to 25 cm or flowers solitary or paired; bracts ±leaflike, divided, not forming involucre. Flowers bisexual, radially symmetric; sepals persistent in fruit [not persistent], 5, yellowish green [white, pink, or purple], plane, ovate to elliptic, 19-30(-50) mm; petals 5-15, distinct, green or brown, funnel-shaped, ± 2-lipped, clawed, 4-8 mm; nectary in center of "funnel"; stamens 30-60; filaments filiform; staminodes absent between stamens and pistils; pistils [2-]3-6[-10], simple, proximally connate [distinct or completely connate]; ovules several per pistil; style present. Fruits follicles [capsules], aggregate, sessile, oblong, sides with prominent transverse veins; beak terminal, straight, 5-15 mm. Seeds usually ± carinate. x = 8.

Distribution

North America, Europe, Asia (in Asia Minor and Tibet).

Discussion

Species ca. 25 (1 in the flora).

Although other species of Helleborus are grown as ornamentals, only the green-flowered H. viridis appears to persist after cultivation. Helleborus niger Linnaeus (Christmas-rose) is a more popular ornamental because of its showy, white to pinkish flowers. It does not appear to persist away from cultivation; it was reported as an escape in 1880 at Sennet, New York, and in 1919 in Washtenaw County, Michigan (R. S. Mitchell and J. K. Dean 1982; E. G. Voss 1972+, vol. 2). Helleborus niger can be distinguished from H. viridis by its flower color and its simple, distal cauline leaves with entire margins.

Both living and dried plants of all species of Helleborus are extremely poisonous. Plants contain a cardiac glycoside (helleborin), which acts directly on the heart muscle, causing convulsions, delirium, and sometimes death. Poisoning from contaminated hay has been known to cause livestock fatalities in some areas (R. S. Mitchell and J. K. Dean 1982).

Selected References

None.

Lower Taxa