Difference between revisions of "Grusonia"

F. Reichenbach ex K. Schumann

Monatsschr. Kakteenk. 6: 177. 1896.

Common names: Club-cholla
Etymology: for Hermann Gruson, 1821–1895, German engineer, and his Magdeburg plant collections
Synonyms: Corynopuntia F. M. KnuthMarenopuntia BackebergMicropuntia DastonOpuntia subg. Corynopuntia (F. M. Knuth) Bra voOpuntia sect. Clavatae EngelmannOpuntia (sect. Undefined) ser. Clavatae (Engelmann) K. Schumann
Treatment appears in FNA Volume 4. Treatment on page 118. Mentioned on page 95.
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--><span class="statement" id="st-d0_s0" data-properties="shrub architecture"><b>Shrubs,</b> usually densely branched, forming mats or clumps [erect or decumbent].</span> <span class="statement" id="st-d0_s1" data-properties="root shape;root density;root derivation"><b>Roots </b>tuberlike and/or diffuse, also often adventitious.</span> <span class="statement" id="st-d0_s2" data-properties="stem segment fixation;stem segment shape;stem segment length;stem segment pubescence;stem segment relief;stem segment architecture or shape;base fragility;base shape;base course"><b>Stem </b>segments firmly attached in most species in the flora to easily dislodged, cylindric-clavate, usually curving upward from near bases, sometimes subspheric, subequal in length, glabrous, strongly tuberculate, sometimes appearing ribbed;</span> <span class="statement" id="st-d0_s3" data-properties="areole arrangement or shape;areole diameter">areoles ± circular, 1–6 mm diam.;</span> <span class="statement" id="st-d0_s4" data-properties="wool coloration;wool coloration;wool coloration;wool coloration;wool coloration">wool white, gray, or yellow.</span> <span class="statement" id="st-d0_s5" data-properties="epidermis sheath duration;spine tip coloration"><b>Spines </b>with epidermis sheath deciduous at apices only, exposing yellow spine tips;</span> <span class="statement" id="st-d0_s6" data-properties="spine count;spine size;distal areole shape;distal areole shape;distal areole shape;distal areole coloration or pubescence or relief;distal areole architecture">at least 1 of major spines in distal areoles angular-flattened to ribbonlike, sometimes cross-striate, sometimes bulbous-based.</span> <span class="statement" id="st-d0_s7" data-properties="glochid coloration;glochid coloration;glochid coloration;glochid coloration;glochid coloration"><b>Glochids </b>commonly increasing greatly in number with time, in distal areoles, white, yellow, or brown.</span> <span class="statement" id="st-d0_s8" data-properties="flower reproduction;flower architecture or shape"><b>Flowers </b>bisexual, radially symmetric;</span> <span class="statement" id="st-d0_s9" data-properties="inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration">inner tepals bright-yellow or rose to purple.</span> <span class="statement" id="st-d0_s10" data-properties="pollen coloration or relief;pollen architecture or coloration or relief"><b>Pollen </b>spinulo-punctate, not reticulate (cylindropuntioid type).</span> <span class="statement" id="st-d0_s11" data-properties="fruit coloration;fruit coloration;fruit coloration;fruit shape;fruit shape;fruit shape;fruit texture;fruit condition;fruit architecture or pubescence or relief;fruit architecture or shape"><b>Fruits </b>yellow to brownish, narrowly obconic to ellipsoid when fertile, fleshy at first, soon drying, smooth, sometimes spiny;</span> <span class="statement" id="st-d0_s12" data-properties="tuft prominence;wool length or size;wool coloration;wool coloration;wool coloration">areoles bearing conspicuous tuft of long, white to tan wool.</span> <span class="statement" id="st-d0_s13" data-properties="seed coloration;seed coloration;seed coloration;seed coloration;seed shape;seed shape;seed some measurement;seed architecture or pubescence or relief;seed architecture or pubescence or relief;bump prominence"><b>Seeds </b>yellowish white or pale-yellow to brownish, suborbicular (to rectangular) or flattened, 3–6 mm, smooth or with rows of raised bumps;</span> <span class="statement" id="st-d0_s14" data-properties="">funicular girdle encircling seeds, not protruding.</span> <span class="statement" id="st-d0_s15" data-properties="girdle derivation;girdle prominence;x chromosome count">x = 11.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Shrubs,</b> usually densely branched, forming mats or clumps [erect or decumbent]. <b>Roots</b> tuberlike and/or diffuse, also often adventitious. <b>Stem</b> segments firmly attached in most species in the flora to easily dislodged, cylindric-clavate, usually curving upward from near bases, sometimes subspheric, subequal in length, glabrous, strongly tuberculate, sometimes appearing ribbed; areoles ± circular, 1–6 mm diam.; wool white, gray, or yellow. <b>Spines</b> with epidermis sheath deciduous at apices only, exposing yellow spine tips; at least 1 of major spines in distal areoles angular-flattened to ribbonlike, sometimes cross-striate, sometimes bulbous based. <b>Glochids</b> commonly increasing greatly in number with time, in distal areoles, white, yellow, or brown. <b>Flowers</b> bisexual, radially symmetric; inner tepals bright yellow or rose to purple. <b>Pollen</b> spinulo-punctate, not reticulate (cylindropuntioid type). <b>Fruits</b> yellow to brownish, narrowly obconic to ellipsoid when fertile, fleshy at first, soon drying, smooth, sometimes spiny; areoles bearing conspicuous tuft of long, white to tan wool. <b>Seeds</b> yellowish white or pale yellow to brownish, suborbicular (to rectangular) or flattened, 3–6 mm, smooth or with rows of raised bumps; funicular girdle encircling seeds, not protruding. <b>x</b> = 11.</span><!--
  
 
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|publication year=1896
 
|publication year=1896
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_229.xml
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V4/V4_229.xml
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|genus=Grusonia
 
|genus=Grusonia
|areole arrangement or shape=circular
 
|areole diameter=1mm;6mm
 
|base course=curving
 
|base fragility=dislodged
 
|base shape=cylindric-clavate
 
|bump prominence=raised
 
|distal areole architecture=bulbous-based
 
|distal areole coloration or pubescence or relief=cross-striate
 
|distal areole shape=angular-flattened;ribbonlike
 
|epidermis sheath duration=deciduous
 
|flower architecture or shape=symmetric
 
|flower reproduction=bisexual
 
|fruit architecture or pubescence or relief=smooth
 
|fruit architecture or shape=spiny
 
|fruit coloration=yellow;brownish
 
|fruit condition=drying
 
|fruit shape=narrowly obconic;ellipsoid
 
|fruit texture=fleshy
 
|girdle derivation=funicular
 
|girdle prominence=not protruding
 
|glochid coloration=brown;yellow;brown;yellow;white
 
|inner tepal coloration=rose;purple
 
|pollen architecture or coloration or relief=not reticulate
 
|pollen coloration or relief=spinulo-punctate
 
|root density=diffuse
 
|root derivation=adventitious
 
|root shape=tuberlike
 
|seed architecture or pubescence or relief=with rows;smooth
 
|seed coloration=pale-yellow;brownish
 
|seed shape=flattened;suborbicular
 
|seed some measurement=3mm;6mm
 
|shrub architecture=branched
 
|spine count=1
 
|spine size=major
 
|spine tip coloration=yellow
 
|stem segment architecture or shape=ribbed
 
|stem segment fixation=attached
 
|stem segment length=subequal
 
|stem segment pubescence=glabrous
 
|stem segment relief=tuberculate
 
|stem segment shape=subspheric
 
|tuft prominence=conspicuous
 
|wool coloration=white;tan
 
|wool length or size=long
 
|x chromosome count=11
 
 
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-->[[Category:Treatment]][[Category:Cactaceae subfam. Opuntioideae]]

Revision as of 15:00, 27 July 2019

Shrubs, usually densely branched, forming mats or clumps [erect or decumbent]. Roots tuberlike and/or diffuse, also often adventitious. Stem segments firmly attached in most species in the flora to easily dislodged, cylindric-clavate, usually curving upward from near bases, sometimes subspheric, subequal in length, glabrous, strongly tuberculate, sometimes appearing ribbed; areoles ± circular, 1–6 mm diam.; wool white, gray, or yellow. Spines with epidermis sheath deciduous at apices only, exposing yellow spine tips; at least 1 of major spines in distal areoles angular-flattened to ribbonlike, sometimes cross-striate, sometimes bulbous based. Glochids commonly increasing greatly in number with time, in distal areoles, white, yellow, or brown. Flowers bisexual, radially symmetric; inner tepals bright yellow or rose to purple. Pollen spinulo-punctate, not reticulate (cylindropuntioid type). Fruits yellow to brownish, narrowly obconic to ellipsoid when fertile, fleshy at first, soon drying, smooth, sometimes spiny; areoles bearing conspicuous tuft of long, white to tan wool. Seeds yellowish white or pale yellow to brownish, suborbicular (to rectangular) or flattened, 3–6 mm, smooth or with rows of raised bumps; funicular girdle encircling seeds, not protruding. x = 11.

Distribution

s United States, n Mexico.

Discussion

Species 14 (8 in the flora).

Key

1 Flowers rose to purple; spines of fruits hairlike, flexible, antrorsely barbed (high magni- fication) Grusonia pulchella
1 Flowers yellow; spines of fruits, when present, more rigid, retrorsely barbed > 2
2 Major adaxial spine(s) of distal areoles terete to subterete or absent; shrubs forming mats or clumps; roots tuberlike > 3
2 Major adaxial spines of distal areoles angular-flattened, at least at bases; shrubs forming mats; roots diffuse > 4
3 Major adaxial spine(s) of distal areoles white to brown to black (but not rich red-brown) or absent; longest central abaxial spine commonly twisted or curved, completely chalky white (at least adaxially) except in basal 3 mm or less Grusonia aggeria
3 Major adaxial spines of distal areoles rich red-brown (or tan); longest central abaxial spine usually straight, brownish, and cross-banded with interrupted, whitish, wavy lines Grusonia grahamii
4 Tubercles very prominent, broad, 1-3.5 times longer than wide, not or little obscured by interlacing spines > 5
4 Tubercles prominent, narrow, 4-6 times longer than wide, obscured by interlacing spines > 6
5 Stem segments 7-19 × 2.5-5 cm; distal tubercles 25-50 mm Grusonia emoryi
5 Stem segments (2-)3.5-6.5 × (1.5-)2-3 cm; distal tubercles 8-20 mm Grusonia schottii
6 Mature fruits very spiny and glochidiate; plants to 50 cm, in series usually 2-4 stem segments. Grusonia kunzei
6 Mature fruits not spiny but glochidiate; plants to 20 cm, in series usually 1-2 stem segments > 7
7 Stem segments 2.5-5(-7.5) cm; distal tubercles 5-10(-16) mm; spines of distal areoles tan or white; longest abaxial spine white, daggerlike, broadly tapered; New Mexico Grusonia clavata
7 Stem segments 5-9 cm; distal tubercles 15-25 mm; spines of distal areoles blackish or brown to tan; longest abaxial spine brown to white, narrowly tapered; Arizona, California, Nevada Grusonia parishii
... more about "Grusonia"
Donald J. Pinkava +
F. Reichenbach ex K. Schumann +
Club-cholla +
s United States +  and n Mexico. +
for Hermann Gruson, 1821–1895, German engineer, and his Magdeburg plant collections +
Monatsschr. Kakteenk. +
pinkava1999b +
Corynopuntia +, Marenopuntia +, Micropuntia +, Opuntia subg. Corynopuntia +, Opuntia sect. Clavatae +  and Opuntia (sect. Undefined) ser. Clavatae +
Grusonia +
Cactaceae subfam. Opuntioideae +