Difference between revisions of "Diphasiastrum tristachyum"

(Pursh) Holub

Preslia 47: 108. 1975.

Common names: Blue ground-cedar lycopode à trois épis
Basionym: Lycopodium tristachyum Pursh Fl. Amer. Sept. 2: 653. 1814
Treatment appears in FNA Volume 2.
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|common_names=Blue ground-cedar;lycopode à trois épis
 
|common_names=Blue ground-cedar;lycopode à trois épis
|basionyms={{Treatment/ID/Synonym
+
|basionyms={{Treatment/ID/Basionym
 
|name=Lycopodium tristachyum
 
|name=Lycopodium tristachyum
 
|authority=Pursh
 
|authority=Pursh
 +
|rank=species
 +
|publication_title=Fl. Amer. Sept.
 +
|publication_place=2: 653. 1814
 
}}
 
}}
 
|synonyms=
 
|synonyms=
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--><span class="statement" id="st-d0_s0" data-properties="stem orientation;stem location;stem width"><b>Horizontal </b>stems deeply (5–12 cm) buried, 1.5–3.2 mm wide;</span> <span class="statement" id="st-d0_s1" data-properties="leaf shape;leaf shape;leaf count;leaf count;apex architecture or relief;apex shape">leaves spatulate to somewhat obovate, 1.8–3.5 X 1.1–1.5, apex faintly erose to irregularly lobed.</span> <span class="statement" id="st-d0_s2" data-properties="shoot orientation;shoot arrangement or growth form;shoot architecture;shoot some measurement"><b>Upright </b>shoots clustered, branching near base, 17–36 cm;</span> <span class="statement" id="st-d0_s3" data-properties="leaf architecture;leaf fixation or orientation;leaf shape;leaf length;leaf width;apex shape">leaves monomorphic, appressed, subulate, 1.9–3.4 X 0.6–1 mm, apex acute.</span> <span class="statement" id="st-d0_s4" data-properties="branchlet shape;branchlet width;angle shape;bud constriction duration;bud constriction shape;bud constriction prominence"><b>Branchlets </b>square with rounded angles in cross-section, 1–2.2 mm wide, annual bud constrictions abrupt and conspicuous;</span> <span class="statement" id="st-d0_s5" data-properties="upperside shape;upperside coloration;upperside coloration;upperside coloration">upperside convex, bluish to whitish green.</span> <span class="statement" id="st-d0_s6" data-properties="branchlet arrangement;upperside leaf fixation or orientation;upperside leaf architecture or shape;portion fusion;portion length;portion width"><b>Leaves </b>on branchlets 4-ranked, upperside leaves appressed, needlelike, free portion of blade 1–1.7 X 0.5–0.9 mm;</span> <span class="statement" id="st-d0_s7" data-properties="lateral leaf fixation or orientation;lateral leaf length;lateral leaf width">lateral leaves appressed, 3.4–7.2 X 1.1–2 mm;</span> <span class="statement" id="st-d0_s8" data-properties="underside leaf fixation or orientation;underside leaf shape;underside leaf length;underside leaf width">underside leaves appressed, somewhat decurrent, 1–2 X 0.4–0.7 mm.</span> <span class="statement" id="st-d0_s9" data-properties="peduncle atypical count;peduncle count;peduncle length;peduncle width"><b>Peduncles </b>(1–) 3, 4–15 X 0.4–1 mm;</span> <span class="statement" id="st-d0_s10" data-properties="leaf arrangement or density;leaf arrangement;leaf shape;tip fusion;tip orientation;tip shape;tip length;tip width">leaves remote, scattered, decurrent, free tips ascending, subulate, 2–3 X 0.2–0.25 mm.</span> <span class="statement" id="st-d0_s11" data-properties="stalk arrangement;peduncle development;peduncle architecture or shape;branch development;branch architecture or shape"><b>Stalks </b>mostly formed by successive forking of peduncle, branches uniformly spaced.</span> <span class="statement" id="st-d0_s12" data-properties="strobilus atypical count;strobilus atypical count;strobilus count;strobilus length;strobilus width;apex architecture;tip reproduction;tip count"><b>Strobili </b>(2–) 3–4 (–7), 10–28 X 2–3 mm, apex round-tipped, sterile tips absent.</span> <span class="statement" id="st-d0_s13" data-properties="sporophyll shape;sporophyll length;sporophyll width"><b>Sporophylls </b>deltate, 2.2–3.5 X 1.6–3 mm, apex gradually tapering.</span> <span class="statement" id="st-d0_s14" data-properties="apex shape;2n chromosome count">2n = 46.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Horizontal </b>stems deeply (5–12 cm) buried, 1.5–3.2 mm wide; leaves spatulate to somewhat obovate, 1.8–3.5 × 1.1–1.5, apex faintly erose to irregularly lobed. <b>Upright</b> shoots clustered, branching near base, 17–36 cm; leaves monomorphic, appressed, subulate, 1.9–3.4 × 0.6–1 mm, apex acute. <b>Branchlets</b> square with rounded angles in cross section, 1–2.2 mm wide, annual bud constrictions abrupt and conspicuous; upperside convex, bluish to whitish green. <b>Leaves</b> on branchlets 4-ranked, upperside leaves appressed, needlelike, free portion of blade 1–1.7 × 0.5–0.9 mm; lateral leaves appressed, 3.4–7.2 × 1.1–2 mm; underside leaves appressed, somewhat decurrent, 1–2 × 0.4–0.7 mm. <b>Peduncles</b> (1–)3, 4–15 × 0.4–1 mm; leaves remote, scattered, decurrent, free tips ascending, subulate, 2–3 × 0.2–0.25 mm. <b>Stalks</b> mostly formed by successive forking of peduncle, branches uniformly spaced. <b>Strobili</b> (2–)3–4(–7), 10–28 × 2–3 mm, apex round-tipped, sterile tips absent. <b>Sporophylls</b> deltate, 2.2–3.5 × 1.6–3 mm, apex gradually tapering. <b>2n</b> = 46.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
 
|habitat=Sterile, acidic soils in open conifer and oak forests
 
|habitat=Sterile, acidic soils in open conifer and oak forests
 
|elevation=50–1800 m
 
|elevation=50–1800 m
|distribution=Man.;N.B.;Nfld.;N.S.;Ont.;P.E.I.;Que.;Ala.;Conn.;Del.;D.C.;Ga.;Ind.;Ky.;Maine;Md.;Mass.;Mich.;Minn.;Mo.;N.H.;N.J.;N.Y.;N.C.;Ohio;Pa.;R.I.;S.C.;Tenn.;Vt.;Va.;W.Va.;Wis.;Europe;Asia in w China.
+
|distribution=Man.;N.B.;Nfld. and Labr. (Nfld.);N.S.;Ont.;P.E.I.;Que.;Ala.;Conn.;Del.;D.C.;Ga.;Ind.;Ky.;Maine;Md.;Mass.;Mich.;Minn.;Mo.;N.H.;N.J.;N.Y.;N.C.;Ohio;Pa.;R.I.;S.C.;Tenn.;Vt.;Va.;W.Va.;Wis.;Europe;Asia in w China.
|discussion=<p>The distinctive Diphasiastrum tristachyum has narrow, rounded branches and dull, bluish white color. It is a parent in more hybrid combinations than any other North American Diphasiastrum.</p><!--
+
|discussion=<p>The distinctive <i>Diphasiastrum tristachyum</i> has narrow, rounded branches and dull, bluish white color. It is a parent in more hybrid combinations than any other North American <i>Diphasiastrum</i>.</p><!--
--><p>The reticulogram shows the known pattern of interspecific hybridization in Diphasiastrum. The hybrids are discussed in detail by J. H. Wilce (1965), and their cytology is summarized by F. S. Wagner (1992). The best known North American hybrids are the four involving D. tristachyum. All of the hybrids have apparently normal meiosis and spores.</p><!--
+
--><p>The reticulogram shows the known pattern of interspecific hybridization in <i>Diphasiastrum</i>. The hybrids are discussed in detail by J. H. Wilce (1965), and their cytology is summarized by F. S. Wagner (1992). The best known North American hybrids are the four involving <i>D. tristachyum</i>. All of the hybrids have apparently normal meiosis and spores.</p><!--
--><p>Diphasiastrum × zeilleri (Rouy) Holub (= D. complanatum X tristachyum) is a frequent plant in areas of distributional overlap between the parents, especially in north central and western Minnesota jackpine forests.</p><!--
+
--><p><i>Diphasiastrum</i> × zeilleri (Rouy) Holub (= <i>D. complanatum</i> X tristachyum) is a frequent plant in areas of distributional overlap between the parents, especially in north central and western Minnesota jackpine forests.</p><!--
--><p>Diphasiastrum × habereri (House) Holub (= D. digitatum X tristachyum) has been overlooked and confused with both parents in zones of overlap. It is found occasionally to frequently in habitats like those of the parents, not necessarily growing close to them.</p><!--
+
--><p><i>Diphasiastrum</i> × habereri (House) Holub (= <i>D. digitatum</i> X tristachyum) has been overlooked and confused with both parents in zones of overlap. It is found occasionally to frequently in habitats like those of the parents, not necessarily growing close to them.</p><!--
--><p>Diphasiastrum × issleri (Rouy) Holub (= D. alpinum X tristachyum) is a rare hybrid in North America, reported only from Maine, but much more widespread in Europe.</p><!--
+
--><p><i>Diphasiastrum</i> × issleri (Rouy) Holub (= <i>D. alpinum</i> X tristachyum) is a rare hybrid in North America, reported only from Maine, but much more widespread in Europe.</p><!--
--><p>Diphasiastrum × sabinifolium (Willdenow) Holub (= D. sitchense X tristachyum) is widespread and frequent in eastern Canada. This hybrid is commonly confused with D. sitchense. It is highly variable, and some individuals approach one or the other parent in morphology (W. J. Cody and D. M. Britton 1989). In the flora, the populations are mainly disjunct from the main range of D. sitchense, including those in Ontario, Quebec, Maine, Michigan, New Hampshire, New York, Pennsylvania, and Vermont.</p><!--
+
--><p><i>Diphasiastrum</i> × sabinifolium (Willdenow) Holub (= <i>D. sitchense</i> X tristachyum) is widespread and frequent in eastern Canada. This hybrid is commonly confused with <i>D. sitchense</i>. It is highly variable, and some individuals approach one or the other parent in morphology (W. J. Cody and D. M. Britton 1989). In the flora, the populations are mainly disjunct from the main range of <i>D. sitchense</i>, including those in Ontario, Quebec, Maine, Michigan, New Hampshire, New York, Pennsylvania, and Vermont.</p><!--
--><p>Two other North American nothospecies are Diphasiastrum complanatum X digitatum and D. alpinum X sitchense.</p>
+
--><p>Two other North American nothospecies are <i>Diphasiastrum complanatum</i> X digitatum and <i>D. alpinum</i> X sitchense.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Diphasiastrum tristachyum
 
name=Diphasiastrum tristachyum
|author=
 
 
|authority=(Pursh) Holub
 
|authority=(Pursh) Holub
 
|rank=species
 
|rank=species
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|habitat=Sterile, acidic soils in open conifer and oak forests
 
|habitat=Sterile, acidic soils in open conifer and oak forests
 
|elevation=50–1800 m
 
|elevation=50–1800 m
|distribution=Man.;N.B.;Nfld.;N.S.;Ont.;P.E.I.;Que.;Ala.;Conn.;Del.;D.C.;Ga.;Ind.;Ky.;Maine;Md.;Mass.;Mich.;Minn.;Mo.;N.H.;N.J.;N.Y.;N.C.;Ohio;Pa.;R.I.;S.C.;Tenn.;Vt.;Va.;W.Va.;Wis.;Europe;Asia in w China.
+
|distribution=Man.;N.B.;Nfld. and Labr. (Nfld.);N.S.;Ont.;P.E.I.;Que.;Ala.;Conn.;Del.;D.C.;Ga.;Ind.;Ky.;Maine;Md.;Mass.;Mich.;Minn.;Mo.;N.H.;N.J.;N.Y.;N.C.;Ohio;Pa.;R.I.;S.C.;Tenn.;Vt.;Va.;W.Va.;Wis.;Europe;Asia in w China.
 
|reference=None
 
|reference=None
 
|publication title=Preslia
 
|publication title=Preslia
 
|publication year=1975
 
|publication year=1975
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_573.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_573.xml
 
|genus=Diphasiastrum
 
|genus=Diphasiastrum
 
|species=Diphasiastrum tristachyum
 
|species=Diphasiastrum tristachyum
|2n chromosome count=46
 
|angle shape=rounded
 
|apex architecture=round-tipped
 
|apex architecture or relief=erose
 
|apex shape=tapering;acute;lobed
 
|branch architecture or shape=forking
 
|branch development=successive
 
|branchlet arrangement=4-ranked
 
|branchlet shape=square
 
|branchlet width=1mm;2.2mm
 
|bud constriction duration=annual
 
|bud constriction prominence=conspicuous
 
|bud constriction shape=abrupt
 
|lateral leaf fixation or orientation=appressed
 
|lateral leaf length=3.4mm;7.2mm
 
|lateral leaf width=1.1mm;2mm
 
|leaf architecture=monomorphic
 
|leaf arrangement=scattered
 
|leaf arrangement or density=remote
 
|leaf count=1.1;1.5
 
|leaf fixation or orientation=appressed
 
|leaf length=1.9mm;3.4mm
 
|leaf shape=decurrent;subulate;obovate;spatulate to somewhat
 
|leaf width=0.6mm;1mm
 
|peduncle architecture or shape=forking
 
|peduncle atypical count=1;3
 
|peduncle count=3
 
|peduncle development=successive
 
|peduncle length=4mm;15mm
 
|peduncle width=0.4mm;1mm
 
|portion fusion=free
 
|portion length=1mm;1.7mm
 
|portion width=0.5mm;0.9mm
 
|shoot architecture=branching
 
|shoot arrangement or growth form=clustered
 
|shoot orientation=upright
 
|shoot some measurement=17cm;36cm
 
|sporophyll length=2.2mm;3.5mm
 
|sporophyll shape=deltate
 
|sporophyll width=1.6mm;3mm
 
|stalk arrangement=spaced
 
|stem location=buried
 
|stem orientation=horizontal
 
|stem width=1.5mm;3.2mm
 
|strobilus atypical count=4;7
 
|strobilus count=3;4
 
|strobilus length=10mm;28mm
 
|strobilus width=2mm;3mm
 
|tip count=absent
 
|tip fusion=free
 
|tip length=2mm;3mm
 
|tip orientation=ascending
 
|tip reproduction=sterile
 
|tip shape=subulate
 
|tip width=0.2mm;0.25mm
 
|underside leaf fixation or orientation=appressed
 
|underside leaf length=1mm;2mm
 
|underside leaf shape=decurrent
 
|underside leaf width=0.4mm;0.7mm
 
|upperside coloration=bluish;whitish green
 
|upperside leaf architecture or shape=needlelike
 
|upperside leaf fixation or orientation=appressed
 
|upperside shape=convex
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Diphasiastrum]]
 
-->[[Category:Treatment]][[Category:Diphasiastrum]]

Latest revision as of 21:24, 5 November 2020

Horizontal stems deeply (5–12 cm) buried, 1.5–3.2 mm wide; leaves spatulate to somewhat obovate, 1.8–3.5 × 1.1–1.5, apex faintly erose to irregularly lobed. Upright shoots clustered, branching near base, 17–36 cm; leaves monomorphic, appressed, subulate, 1.9–3.4 × 0.6–1 mm, apex acute. Branchlets square with rounded angles in cross section, 1–2.2 mm wide, annual bud constrictions abrupt and conspicuous; upperside convex, bluish to whitish green. Leaves on branchlets 4-ranked, upperside leaves appressed, needlelike, free portion of blade 1–1.7 × 0.5–0.9 mm; lateral leaves appressed, 3.4–7.2 × 1.1–2 mm; underside leaves appressed, somewhat decurrent, 1–2 × 0.4–0.7 mm. Peduncles (1–)3, 4–15 × 0.4–1 mm; leaves remote, scattered, decurrent, free tips ascending, subulate, 2–3 × 0.2–0.25 mm. Stalks mostly formed by successive forking of peduncle, branches uniformly spaced. Strobili (2–)3–4(–7), 10–28 × 2–3 mm, apex round-tipped, sterile tips absent. Sporophylls deltate, 2.2–3.5 × 1.6–3 mm, apex gradually tapering. 2n = 46.


Habitat: Sterile, acidic soils in open conifer and oak forests
Elevation: 50–1800 m

Distribution

V2 573-distribution-map.gif

Man., N.B., Nfld. and Labr. (Nfld.), N.S., Ont., P.E.I., Que., Ala., Conn., Del., D.C., Ga., Ind., Ky., Maine, Md., Mass., Mich., Minn., Mo., N.H., N.J., N.Y., N.C., Ohio, Pa., R.I., S.C., Tenn., Vt., Va., W.Va., Wis., Europe, Asia in w China.

Discussion

The distinctive Diphasiastrum tristachyum has narrow, rounded branches and dull, bluish white color. It is a parent in more hybrid combinations than any other North American Diphasiastrum.

The reticulogram shows the known pattern of interspecific hybridization in Diphasiastrum. The hybrids are discussed in detail by J. H. Wilce (1965), and their cytology is summarized by F. S. Wagner (1992). The best known North American hybrids are the four involving D. tristachyum. All of the hybrids have apparently normal meiosis and spores.

Diphasiastrum × zeilleri (Rouy) Holub (= D. complanatum X tristachyum) is a frequent plant in areas of distributional overlap between the parents, especially in north central and western Minnesota jackpine forests.

Diphasiastrum × habereri (House) Holub (= D. digitatum X tristachyum) has been overlooked and confused with both parents in zones of overlap. It is found occasionally to frequently in habitats like those of the parents, not necessarily growing close to them.

Diphasiastrum × issleri (Rouy) Holub (= D. alpinum X tristachyum) is a rare hybrid in North America, reported only from Maine, but much more widespread in Europe.

Diphasiastrum × sabinifolium (Willdenow) Holub (= D. sitchense X tristachyum) is widespread and frequent in eastern Canada. This hybrid is commonly confused with D. sitchense. It is highly variable, and some individuals approach one or the other parent in morphology (W. J. Cody and D. M. Britton 1989). In the flora, the populations are mainly disjunct from the main range of D. sitchense, including those in Ontario, Quebec, Maine, Michigan, New Hampshire, New York, Pennsylvania, and Vermont.

Two other North American nothospecies are Diphasiastrum complanatum X digitatum and D. alpinum X sitchense.

Selected References

None.

Lower Taxa

None.
... more about "Diphasiastrum tristachyum"
Warren H. Wagner Jr. +  and Joseph M. Beitel +
(Pursh) Holub +
Lycopodium tristachyum +
Blue ground-cedar +  and lycopode à trois épis +
Man. +, N.B. +, Nfld. and Labr. (Nfld.) +, N.S. +, Ont. +, P.E.I. +, Que. +, Ala. +, Conn. +, Del. +, D.C. +, Ga. +, Ind. +, Ky. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Mo. +, N.H. +, N.J. +, N.Y. +, N.C. +, Ohio +, Pa. +, R.I. +, S.C. +, Tenn. +, Vt. +, Va. +, W.Va. +, Wis. +, Europe +  and Asia in w China. +
50–1800 m +
Sterile, acidic soils in open conifer and oak forests +
Diphasiastrum tristachyum +
Diphasiastrum +
species +