Difference between revisions of "Cylindropuntia versicolor"

(Engelmann ex J. M. Coulter) F. M. Knuth

in C. Backeberg and F. M. Knuth, Kaktus-ABC, 125. 1935.

Common names: Staghorn cholla
Basionym: Opuntia versicolor Engelmann ex J. M. Coulter Contr. U.S. Natl. Herb. 3: 452. 1896
Synonyms: Opuntia arborescens var. versicolor (Engelmann ex J. M. Coulter) Dams Opuntia thurberi subsp. versicolor (Engelmann ex J. M. Coulter) Felger & Lowe
Treatment appears in FNA Volume 4. Treatment on page 109. Mentioned on page 105, 106, 107, 112, 116.
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{{Treatment/ID
 
{{Treatment/ID
 
|accepted_name=Cylindropuntia versicolor
 
|accepted_name=Cylindropuntia versicolor
|accepted_authority=(Engelmann ex J. M. Coulter) F. M. Knuth in C. Backeberg and F. M. Knuth
+
|accepted_authority=(Engelmann ex J. M. Coulter) F. M. Knuth
 
|publications={{Treatment/Publication
 
|publications={{Treatment/Publication
 
|title=in C. Backeberg and F. M. Knuth, Kaktus-ABC,
 
|title=in C. Backeberg and F. M. Knuth, Kaktus-ABC,
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}}
 
}}
 
|common_names=Staghorn cholla
 
|common_names=Staghorn cholla
|basionyms={{Treatment/ID/Synonym
+
|basionyms={{Treatment/ID/Basionym
 
|name=Opuntia versicolor
 
|name=Opuntia versicolor
 
|authority=Engelmann ex J. M. Coulter
 
|authority=Engelmann ex J. M. Coulter
 +
|rank=species
 +
|publication_title=Contr. U.S. Natl. Herb.
 +
|publication_place=3: 452. 1896
 
}}
 
}}
 
|synonyms={{Treatment/ID/Synonym
 
|synonyms={{Treatment/ID/Synonym
 
|name=Opuntia arborescens var. versicolor
 
|name=Opuntia arborescens var. versicolor
 
|authority=(Engelmann ex J. M. Coulter) Dams
 
|authority=(Engelmann ex J. M. Coulter) Dams
}}{{Treatment/ID/Synonym
+
|rank=variety
 +
}} {{Treatment/ID/Synonym
 
|name=Opuntia thurberi subsp. versicolor
 
|name=Opuntia thurberi subsp. versicolor
 
|authority=(Engelmann ex J. M. Coulter) Felger & Lowe
 
|authority=(Engelmann ex J. M. Coulter) Felger & Lowe
 +
|rank=subspecies
 
}}
 
}}
 
|hierarchy=Cactaceae;Cactaceae subfam. Opuntioideae;Cylindropuntia;Cylindropuntia versicolor
 
|hierarchy=Cactaceae;Cactaceae subfam. Opuntioideae;Cylindropuntia;Cylindropuntia versicolor
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}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="tree architecture;tree some measurement;shrub architecture;shrub some measurement;angle shape"><b>Trees </b>or shrubs, openly branching at acute angles, to 2 m.</span> <span class="statement" id="st-d0_s1" data-properties="stem segment arrangement;stem segment arrangement;stem segment coloration or density;stem segment coloration or density;stem segment coloration or density;stem segment length;stem segment width"><b>Stem </b>segments whorled or subwhorled, purple to green-purple, 4–18 × 1–2 cm;</span> <span class="statement" id="st-d0_s2" data-properties="tubercle prominence;tubercle shape;tubercle atypical some measurement;tubercle some measurement">tubercles prominent, elongate-oval, 1–2 (–2.5) cm;</span> <span class="statement" id="st-d0_s3" data-properties="areole shape;areole diameter">areoles subcircular, 3.5–4 mm diam.;</span> <span class="statement" id="st-d0_s4" data-properties="wool coloration;wool coloration;wool coloration;wool life cycle;wool coloration">wool tan to brown, aging gray.</span> <span class="statement" id="st-d0_s5" data-properties="spine count;spine arrangement;spine arrangement;spine height or length or size;areole arrangement"><b>Spines </b>6–8 per areole (1–2 bristlelike spines) increasing with time, well distributed along stem, slightly interlacing with spines of adjacent areoles, short;</span> <span class="statement" id="st-d0_s6" data-properties="abaxial spine orientation;abaxial spine coloration;abaxial spine coloration;abaxial spine coloration;abaxial spine coloration;abaxial spine shape;abaxial spine length;abaxial spine some measurement">abaxial spines usually reflexed, whitish, pinkish to redbrown, lightly flattened, the longest 10–18 mm;</span> <span class="statement" id="st-d0_s7" data-properties="adaxial spine orientation;adaxial spine orientation;adaxial spine count;adaxial spine coloration;adaxial spine coloration;adaxial spine position relational;adaxial spine shape;adaxial spine some measurement">adaxial spines erect or spreading, rich redbrown, gray coated basally, subterete, 6–11 mm;</span> <span class="statement" id="st-d0_s8" data-properties="sheath coloration;sheath coloration;sheath coloration;sheath coloration;sheath coloration;glochid shape;glochid coloration;glochid some measurement;adaxial tuft size">sheaths grayish, tipped brown or yellow to golden Glochids in small adaxial tuft or crescent, dark yellow, to 1 mm.</span> <span class="statement" id="st-d0_s9" data-properties="inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal shape;inner tepal architecture or shape"><b>Flowers:</b> inner tepals yellow-green, yellow to gold or bronze, or red to rose or magenta, spatulate, apiculate;</span> <span class="statement" id="st-d0_s10" data-properties="filament coloration">filaments yellowish green;</span> <span class="statement" id="st-d0_s11" data-properties="anther coloration">anthers yellow;</span> <span class="statement" id="st-d0_s12" data-properties="style coloration;style coloration;style coloration">style whitish to pale bronze;</span> <span class="statement" id="st-d0_s13" data-properties="stigma lobe coloration">stigma lobes whitish.</span> <span class="statement" id="st-d0_s14" data-properties="fruit life cycle;fruit coloration;fruit coloration;fruit coloration;fruit coloration;fruit architecture;fruit shape;fruit length;fruit width;fruit texture;fruit relief;fruit architecture or pubescence or relief;fruit architecture"><b>Fruits </b>often proliferating, yellowish green or tinged red to purple, often stipitate, obovate, 25–40 × 10–20 mm, leathery-fleshy, tuberculate or smooth, essentially spineless;</span> <span class="statement" id="st-d0_s15" data-properties="tubercle prominence;tubercle size;tubercle reproduction;tubercle size;tubercle architecture or pubescence or relief;proximal tubercle character;proximal tubercle length or size">tubercles prominent, at first, subequal in length or proximal ones longer, fertile ones swelling and becoming smooth;</span> <span class="statement" id="st-d0_s16" data-properties="umbilicus some measurement;umbilicus depth">umbilicus to 9 mm deep;</span> <span class="statement" id="st-d0_s17" data-properties="areole count">areoles 20–30.</span> <span class="statement" id="st-d0_s18" data-properties="seed coloration;seed shape;seed shape;seed length;seed width;side shape;side shape;protrusion arrangement or shape"><b>Seeds </b>yellowish, polygonal in outline, warped, 3.5–5.5 × 2.5–5 mm, sides nearly flattened with depressions and protrusions to angular subspheric;</span> <span class="statement" id="st-d0_s19" data-properties="">girdle smooth or in a groove.</span> <span class="statement" id="st-d0_s20" data-properties="girdle architecture or pubescence or relief;girdle architecture or pubescence or relief;2n chromosome count">2n = 22.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Trees </b>or shrubs, openly branching at acute angles, to 2 m. <b>Stem</b> segments whorled or subwhorled, purple to green-purple, 4–18 × 1–2 cm; tubercles prominent, elongate-oval, 1–2(–2.5) cm; areoles subcircular, 3.5–4 mm diam.; wool tan to brown, aging gray. <b>Spines</b> 6–8 per areole (1–2 bristlelike spines) increasing with time, well distributed along stem, slightly interlacing with spines of adjacent areoles, short; abaxial spines usually reflexed, whitish, pinkish to red-brown, lightly flattened, the longest 10–18 mm; adaxial spines erect or spreading, rich red-brown, gray coated basally, subterete, 6–11 mm; sheaths grayish, tipped brown or yellow to golden Glochids in small adaxial tuft or crescent, dark yellow, to 1 mm. <b>Flowers</b>: inner tepals yellow-green, yellow to gold or bronze, or red to rose or magenta, spatulate, apiculate; filaments yellowish green; anthers yellow; style whitish to pale bronze; stigma lobes whitish. <b>Fruits</b> often proliferating, yellowish green or tinged red to purple, often stipitate, obovate, 25–40 × 10–20 mm, leathery-fleshy, tuberculate or smooth, essentially spineless; tubercles prominent, at first, subequal in length or proximal ones longer, fertile ones swelling and becoming smooth; umbilicus to 9 mm deep; areoles 20–30. <b>Seeds</b> yellowish, polygonal in outline, warped, 3.5–5.5 × 2.5–5 mm, sides nearly flattened with depressions and protrusions to angular subspheric; girdle smooth or in a groove. <b>2n</b> = 22.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
 +
|phenology=Flowering spring (Apr–Jun).
 
|habitat=Sonoran Desert, desert scrub, flats, washes, rocky hillsides, canyons
 
|habitat=Sonoran Desert, desert scrub, flats, washes, rocky hillsides, canyons
 
|elevation=600-1300 m
 
|elevation=600-1300 m
 
|distribution=Ariz.;Mexico (Sonora).
 
|distribution=Ariz.;Mexico (Sonora).
|discussion=<p>Cylindropuntia versicolor forms hybrids with C. acanthocarpa var. major, C. arbuscula (= C. ×vivipara), C. leptocaulis (see discussion under 3. C. ×tetracantha), and C. spinosior (= C. ×grantiorum P. V. Heath). Cylindropuntia ×grantiorum, found in eastern to south-central Arizona, is intermediate between the parental species in values for the number of spines per stem areole, stem segment diameter and color, and fruit tubercles. The hybrids between C. versicolor and C. acanthocarpa var. major, which occur in south-central Arizona, have fleshy, tuberculate fruits, often with one or two short permanent spines at apex and longer and more numerous spines per stem areole.</p><!--
+
|discussion=<p><i>Cylindropuntia versicolor</i> forms hybrids with <i>C. acanthocarpa </i>var.<i> major</i>, <i>C. arbuscula</i> (= C. ×vivipara), <i>C. leptocaulis</i> (see discussion under 3. <i>C. ×tetracantha</i>), and <i>C. spinosior</i> (= C. ×grantiorum P. V. Heath). <i>Cylindropuntia</i> ×grantiorum, found in eastern to south-central Arizona, is intermediate between the parental species in values for the number of spines per stem areole, stem segment diameter and color, and fruit tubercles. The hybrids between <i>C. versicolor</i> and <i>C. acanthocarpa </i>var.<i> major</i>, which occur in south-central Arizona, have fleshy, tuberculate fruits, often with one or two short permanent spines at apex and longer and more numerous spines per stem areole.</p><!--
--><p>Cylindropuntia versicolor itself may be of hybrid origin.</p>
+
--><p><i>Cylindropuntia versicolor</i> itself may be of hybrid origin.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Cylindropuntia versicolor
 
name=Cylindropuntia versicolor
|author=
+
|authority=(Engelmann ex J. M. Coulter) F. M. Knuth
|authority=(Engelmann ex J. M. Coulter) F. M. Knuth in C. Backeberg and F. M. Knuth
 
 
|rank=species
 
|rank=species
 
|parent rank=genus
 
|parent rank=genus
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|basionyms=Opuntia versicolor
 
|basionyms=Opuntia versicolor
 
|family=Cactaceae
 
|family=Cactaceae
 +
|phenology=Flowering spring (Apr–Jun).
 
|habitat=Sonoran Desert, desert scrub, flats, washes, rocky hillsides, canyons
 
|habitat=Sonoran Desert, desert scrub, flats, washes, rocky hillsides, canyons
 
|elevation=600-1300 m
 
|elevation=600-1300 m
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|publication year=1935
 
|publication year=1935
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_205.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V4/V4_205.xml
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|genus=Cylindropuntia
 
|genus=Cylindropuntia
 
|species=Cylindropuntia versicolor
 
|species=Cylindropuntia versicolor
|2n chromosome count=22
 
|abaxial spine coloration=pinkish;redbrown
 
|abaxial spine length=longest
 
|abaxial spine orientation=reflexed
 
|abaxial spine shape=flattened
 
|abaxial spine some measurement=10mm;18mm
 
|adaxial spine coloration=gray;redbrown
 
|adaxial spine count=rich
 
|adaxial spine orientation=spreading;erect
 
|adaxial spine position relational=coated
 
|adaxial spine shape=subterete
 
|adaxial spine some measurement=6mm;11mm
 
|adaxial tuft size=small
 
|angle shape=acute
 
|anther coloration=yellow
 
|areole arrangement=adjacent
 
|areole count=20;30
 
|areole diameter=3.5mm;4mm
 
|areole shape=subcircular
 
|filament coloration=yellowish green
 
|fruit architecture=spineless;stipitate
 
|fruit architecture or pubescence or relief=smooth
 
|fruit coloration=tinged red;purple
 
|fruit length=25mm;40mm
 
|fruit life cycle=proliferating
 
|fruit relief=tuberculate
 
|fruit shape=obovate
 
|fruit texture=leathery-fleshy
 
|fruit width=10mm;20mm
 
|girdle architecture or pubescence or relief=in a groove;smooth
 
|glochid coloration=dark yellow
 
|glochid shape=crescent
 
|glochid some measurement=0mm;1mm
 
|inner tepal architecture or shape=apiculate
 
|inner tepal coloration=yellow;gold or bronze or red
 
|inner tepal shape=spatulate
 
|protrusion arrangement or shape=angular
 
|proximal tubercle character=length
 
|proximal tubercle length or size=longer
 
|seed coloration=yellowish
 
|seed length=3.5mm;5.5mm
 
|seed shape=warped;polygonal
 
|seed width=2.5mm;5mm
 
|sheath coloration=yellow;golden
 
|shrub architecture=branching
 
|shrub some measurement=0m;2m
 
|side shape=subspheric;flattened
 
|spine arrangement=interlacing;distributed
 
|spine count=6;8
 
|spine height or length or size=short
 
|stem segment arrangement=subwhorled;whorled
 
|stem segment coloration or density=purple;green-purple
 
|stem segment length=4cm;18cm
 
|stem segment width=1cm;2cm
 
|stigma lobe coloration=whitish
 
|style coloration=whitish;pale bronze
 
|tree architecture=branching
 
|tree some measurement=0m;2m
 
|tubercle architecture or pubescence or relief=smooth
 
|tubercle atypical some measurement=2cm;2.5cm
 
|tubercle prominence=prominent;prominent
 
|tubercle reproduction=fertile
 
|tubercle shape=elongate-oval
 
|tubercle size=swelling;subequal
 
|tubercle some measurement=1cm;2cm
 
|umbilicus depth=deep
 
|umbilicus some measurement=0mm;9mm
 
|wool coloration=gray;tan;brown
 
|wool life cycle=aging
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Cylindropuntia]]
 
-->[[Category:Treatment]][[Category:Cylindropuntia]]

Latest revision as of 22:57, 5 November 2020

Trees or shrubs, openly branching at acute angles, to 2 m. Stem segments whorled or subwhorled, purple to green-purple, 4–18 × 1–2 cm; tubercles prominent, elongate-oval, 1–2(–2.5) cm; areoles subcircular, 3.5–4 mm diam.; wool tan to brown, aging gray. Spines 6–8 per areole (1–2 bristlelike spines) increasing with time, well distributed along stem, slightly interlacing with spines of adjacent areoles, short; abaxial spines usually reflexed, whitish, pinkish to red-brown, lightly flattened, the longest 10–18 mm; adaxial spines erect or spreading, rich red-brown, gray coated basally, subterete, 6–11 mm; sheaths grayish, tipped brown or yellow to golden Glochids in small adaxial tuft or crescent, dark yellow, to 1 mm. Flowers: inner tepals yellow-green, yellow to gold or bronze, or red to rose or magenta, spatulate, apiculate; filaments yellowish green; anthers yellow; style whitish to pale bronze; stigma lobes whitish. Fruits often proliferating, yellowish green or tinged red to purple, often stipitate, obovate, 25–40 × 10–20 mm, leathery-fleshy, tuberculate or smooth, essentially spineless; tubercles prominent, at first, subequal in length or proximal ones longer, fertile ones swelling and becoming smooth; umbilicus to 9 mm deep; areoles 20–30. Seeds yellowish, polygonal in outline, warped, 3.5–5.5 × 2.5–5 mm, sides nearly flattened with depressions and protrusions to angular subspheric; girdle smooth or in a groove. 2n = 22.


Phenology: Flowering spring (Apr–Jun).
Habitat: Sonoran Desert, desert scrub, flats, washes, rocky hillsides, canyons
Elevation: 600-1300 m

Discussion

Cylindropuntia versicolor forms hybrids with C. acanthocarpa var. major, C. arbuscula (= C. ×vivipara), C. leptocaulis (see discussion under 3. C. ×tetracantha), and C. spinosior (= C. ×grantiorum P. V. Heath). Cylindropuntia ×grantiorum, found in eastern to south-central Arizona, is intermediate between the parental species in values for the number of spines per stem areole, stem segment diameter and color, and fruit tubercles. The hybrids between C. versicolor and C. acanthocarpa var. major, which occur in south-central Arizona, have fleshy, tuberculate fruits, often with one or two short permanent spines at apex and longer and more numerous spines per stem areole.

Cylindropuntia versicolor itself may be of hybrid origin.

Selected References

None.

Lower Taxa

None.
... more about "Cylindropuntia versicolor"
Donald J. Pinkava +
(Engelmann ex J. M. Coulter) F. M. Knuth +
Opuntia versicolor +
Staghorn cholla +
Ariz. +  and Mexico (Sonora). +
600-1300 m +
Sonoran Desert, desert scrub, flats, washes, rocky hillsides, canyons +
Flowering spring (Apr–Jun). +
in C. Backeberg and F. M. Knuth, Kaktus-ABC, +
Opuntia arborescens var. versicolor +  and Opuntia thurberi subsp. versicolor +
Cylindropuntia versicolor +
Cylindropuntia +
species +