Difference between revisions of "Cylindropuntia acanthocarpa"

(Engelmann & J. M. Bigelow) F. M. Knuth

in C. Backeberg and F. M. Knuth, Kaktus-ABC, 124. 1935.

Common names: Buckhorn cholla
Illustrated
Basionym: Opuntia acanthocarpa Engelmann & J. M. Bigelow Proc. Amer. Acad. Arts 3: 308. 1856
Treatment appears in FNA Volume 4. Mentioned on page 104, 107, 110, 111.
FNA>Volume Importer
 
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{{Treatment/ID
 
{{Treatment/ID
 
|accepted_name=Cylindropuntia acanthocarpa
 
|accepted_name=Cylindropuntia acanthocarpa
|accepted_authority=(Engelmann & J. M. Bigelow) F. M. Knuth in C. Backeberg and F. M. Knuth
+
|accepted_authority=(Engelmann & J. M. Bigelow) F. M. Knuth
 
|publications={{Treatment/Publication
 
|publications={{Treatment/Publication
 
|title=in C. Backeberg and F. M. Knuth, Kaktus-ABC,
 
|title=in C. Backeberg and F. M. Knuth, Kaktus-ABC,
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}}
 
}}
 
|common_names=Buckhorn cholla
 
|common_names=Buckhorn cholla
|basionyms={{Treatment/ID/Synonym
+
|special_status={{Treatment/ID/Special_status
 +
|code=F
 +
|label=Illustrated
 +
}}
 +
|basionyms={{Treatment/ID/Basionym
 
|name=Opuntia acanthocarpa
 
|name=Opuntia acanthocarpa
 
|authority=Engelmann & J. M. Bigelow
 
|authority=Engelmann & J. M. Bigelow
 +
|rank=species
 +
|publication_title=Proc. Amer. Acad. Arts
 +
|publication_place=3: 308. 1856
 
}}
 
}}
 
|synonyms=
 
|synonyms=
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}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="tree architecture;tree atypical some measurement;tree some measurement;shrub architecture;shrub atypical some measurement;shrub some measurement"><b>Trees </b>or shrubs, sparsely to densely branched, 1–2 (–4) m.</span> <span class="statement" id="st-d0_s1" data-properties="stem segment fixation;stem segment shape;stem segment atypical length;stem segment length;stem segment atypical width;stem segment width"><b>Stem </b>segments firmly attached, cylindric, 10–30 (–50) × 2–2.5 (–3) cm;</span> <span class="statement" id="st-d0_s2" data-properties="tubercle prominence;tubercle width;tubercle width;tubercle width;tubercle some measurement">tubercles prominent, narrow to broad, 1.5–4.5 cm;</span> <span class="statement" id="st-d0_s3" data-properties="areole shape;areole shape;areole shape;areole length;areole width">areoles elliptic to subcircular, 4–5.5 × 4–5 mm;</span> <span class="statement" id="st-d0_s4" data-properties="wool coloration;wool coloration;wool coloration;wool coloration;wool life cycle;wool coloration">wool white or yellowish to tan, aging gray-black.</span> <span class="statement" id="st-d0_s5" data-properties="spine atypical count;spine count;spine count;spine height or length or size;spine architecture;areole margin count;areole margin height or length or size;areole margin architecture"><b>Spines </b>6–20 (–30+) per areole, often accompanied by 0–5 short bristlelike spines at areole margins, usually in distal areoles;</span> <span class="statement" id="st-d0_s6" data-properties="abaxial spine size;abaxial spine orientation;abaxial spine arrangement;abaxial spine shape;abaxial spine shape;abaxial spine shape">major abaxial spines deflexed to divergent, subterete to flattened;</span> <span class="statement" id="st-d0_s7" data-properties="adaxial spine size;adaxial spine arrangement;adaxial spine shape;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine life cycle;adaxial spine coloration;central spine length;central spine atypical some measurement;central spine some measurement">major adaxial spines ascending-divergent, terete, yellow or tan to deep redbrown, aging gray, the central spine longest, 12–30 (–38) mm;</span> <span class="statement" id="st-d0_s8" data-properties="sheath coloration;sheath coloration;sheath coloration;sheath size;tip coloration;tip coloration;tip coloration">sheaths white to grayish with yellow to golden tips, baggy to tight-fitting.</span> <span class="statement" id="st-d0_s9" data-properties="glochid coloration;glochid coloration;glochid coloration;glochid some measurement;adaxial tuft prominence;adaxial tuft size"><b>Glochids </b>in inconspicuous to small adaxial tuft, yellow to brown, 0.5–2 mm.</span> <span class="statement" id="st-d0_s10" data-properties="inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal shape;inner tepal some measurement;inner tepal architecture or shape"><b>Flowers:</b> inner tepals bright-yellow to bronze to brick-red, spatulate, 20–30 mm, apiculate-emarginate;</span> <span class="statement" id="st-d0_s11" data-properties="filament coloration">filaments red;</span> <span class="statement" id="st-d0_s12" data-properties="anther coloration">anthers yellow;</span> <span class="statement" id="st-d0_s13" data-properties="style coloration;style coloration;style coloration;stigma lobe coloration;stigma lobe coloration;stigma lobe coloration">style and stigma lobes white to light green.</span> <span class="statement" id="st-d0_s14" data-properties="fruit coloration;fruit shape;fruit shape;fruit shape;fruit length;fruit width;fruit condition or texture;fruit relief;fruit architecture or shape;umbilicus depth"><b>Fruits </b>tan at maturity, obconic to ellipsoid, 15–35 × 15–20 mm, dry, tuberculate, densely to sparsely spiny (rarely spineless), with apical flange above shallow umbilicus;</span> <span class="statement" id="st-d0_s15" data-properties="basal tubercle length">basal tubercles longest;</span> <span class="statement" id="st-d0_s16" data-properties="areole count">areoles 12–30.</span> <span class="statement" id="st-d0_s17" data-properties="seed coloration;seed coloration;seed coloration;seed shape;seed shape;seed shape;seed length;seed width;side architecture or pubescence or relief;depression count;depression size"><b>Seeds </b>pale-yellow to tan, angular or squarish in outline, warped, 3.5–5 × 3–4.5 mm, sides smooth, each with 2–4 large depressions;</span> <span class="statement" id="st-d0_s18" data-properties="girdle architecture or pubescence or relief">girdle smooth.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Trees </b>or shrubs, sparsely to densely branched, 1–2(–4) m. <b>Stem</b> segments firmly attached, cylindric, 10–30(–50) × 2–2.5(–3) cm; tubercles prominent, narrow to broad, 1.5–4.5 cm; areoles elliptic to subcircular, 4–5.5 × 4–5 mm; wool white or yellowish to tan, aging gray-black. <b>Spines</b> 6–20(–30+) per areole, often accompanied by 0–5 short bristlelike spines at areole margins, usually in distal areoles; major abaxial spines deflexed to divergent, subterete to flattened; major adaxial spines ascending-divergent, terete, yellow or tan to deep red-brown, aging gray, the central spine longest, 12–30(–38) mm; sheaths white to grayish with yellow to golden tips, baggy to tight fitting. <b>Glochids</b> in inconspicuous to small adaxial tuft, yellow to brown, 0.5–2 mm. <b>Flowers</b>: inner tepals bright yellow to bronze to brick red, spatulate, 20–30 mm, apiculate-emarginate; filaments red; anthers yellow; style and stigma lobes white to light green. <b>Fruits</b> tan at maturity, obconic to ellipsoid, 15–35 × 15–20 mm, dry, tuberculate, densely to sparsely spiny (rarely spineless), with apical flange above shallow umbilicus; basal tubercles longest; areoles 12–30. <b>Seeds</b> pale yellow to tan, angular or squarish in outline, warped, 3.5–5 × 3–4.5 mm, sides smooth, each with 2–4 large depressions; girdle smooth.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
|distribution=sw United States;n Mexico.
+
|distribution=Ariz.;Calif.;Nev.;Utah;n Mexico.
 
|discussion=<p>Varieties 4 (4 in the flora).</p><!--
 
|discussion=<p>Varieties 4 (4 in the flora).</p><!--
--><p>Cylindropuntia acanthocarpa hybridizes with C. abyssi, C. echinocarpa (= C. ×deserta), C. bigelovii (= C. ×campii), C. leptocaulis (see C. ×tetracantha), C. ×multigeniculata, C. spinosior, C. versicolor, and C. whipplei (= C. congesta). Hybrids of C. acanthocarpa and C. ×multigeniculata are more open in habit, with longer stem segments than C. ×multigeniculata and with reddish filaments like C. acanthocarpa.</p>
+
--><p><i>Cylindropuntia acanthocarpa</i> hybridizes with <i>C. abyssi</i>, <i>C. echinocarpa</i> (= C. ×deserta), <i>C. bigelovii</i> (= C. ×campii), <i>C. leptocaulis</i> (see <i>C. ×tetracantha</i>), C. ×multigeniculata, <i>C. spinosior</i>, <i>C. versicolor</i>, and <i>C. whipplei</i> (= C. congesta). Hybrids of <i>C. acanthocarpa</i> and C. ×multigeniculata are more open in habit, with longer stem segments than C. ×multigeniculata and with reddish filaments like <i>C. acanthocarpa</i>.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Cylindropuntia acanthocarpa
 
name=Cylindropuntia acanthocarpa
|author=
+
|authority=(Engelmann & J. M. Bigelow) F. M. Knuth
|authority=(Engelmann & J. M. Bigelow) F. M. Knuth in C. Backeberg and F. M. Knuth
 
 
|rank=species
 
|rank=species
 
|parent rank=genus
 
|parent rank=genus
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|basionyms=Opuntia acanthocarpa
 
|basionyms=Opuntia acanthocarpa
 
|family=Cactaceae
 
|family=Cactaceae
|distribution=sw United States;n Mexico.
+
|distribution=Ariz.;Calif.;Nev.;Utah;n Mexico.
 
|reference=None
 
|reference=None
 
|publication title=in C. Backeberg and F. M. Knuth, Kaktus-ABC,
 
|publication title=in C. Backeberg and F. M. Knuth, Kaktus-ABC,
 
|publication year=1935
 
|publication year=1935
|special status=
+
|special status=Illustrated
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_222.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V4/V4_222.xml
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|genus=Cylindropuntia
 
|genus=Cylindropuntia
 
|species=Cylindropuntia acanthocarpa
 
|species=Cylindropuntia acanthocarpa
|abaxial spine arrangement=divergent
 
|abaxial spine orientation=deflexed
 
|abaxial spine shape=subterete;flattened
 
|abaxial spine size=major
 
|adaxial spine arrangement=ascending-divergent
 
|adaxial spine coloration=gray;tan;deep redbrown
 
|adaxial spine life cycle=aging
 
|adaxial spine shape=terete
 
|adaxial spine size=major
 
|adaxial tuft prominence=inconspicuous
 
|adaxial tuft size=small
 
|anther coloration=yellow
 
|areole count=12;30
 
|areole length=4mm;5.5mm
 
|areole margin architecture=bristlelike
 
|areole margin count=0;5
 
|areole margin height or length or size=short
 
|areole shape=elliptic;subcircular
 
|areole width=4mm;5mm
 
|basal tubercle length=longest
 
|central spine atypical some measurement=30mm;38mm
 
|central spine length=longest
 
|central spine some measurement=12mm;30mm
 
|depression count=2;4
 
|depression size=large
 
|filament coloration=red
 
|fruit architecture or shape=spiny
 
|fruit coloration=tan
 
|fruit condition or texture=dry
 
|fruit length=15mm;35mm
 
|fruit relief=tuberculate
 
|fruit shape=obconic;ellipsoid
 
|fruit width=15mm;20mm
 
|girdle architecture or pubescence or relief=smooth
 
|glochid coloration=yellow;brown
 
|glochid some measurement=0.5mm;2mm
 
|inner tepal architecture or shape=apiculate-emarginate
 
|inner tepal coloration=bright-yellow;bronze
 
|inner tepal shape=spatulate
 
|inner tepal some measurement=20mm;30mm
 
|seed coloration=pale-yellow;tan
 
|seed length=3.5mm;5mm
 
|seed shape=warped;squarish;angular
 
|seed width=3mm;4.5mm
 
|sheath coloration=white;grayish
 
|sheath size=tight-fitting
 
|shrub architecture=branched
 
|shrub atypical some measurement=2m;4m
 
|shrub some measurement=1m;2m
 
|side architecture or pubescence or relief=smooth
 
|spine architecture=bristlelike
 
|spine atypical count=20;30
 
|spine count=0;5
 
|spine height or length or size=short
 
|stem segment atypical length=30cm;50cm
 
|stem segment atypical width=2.5cm;3cm
 
|stem segment fixation=attached
 
|stem segment length=10cm;30cm
 
|stem segment shape=cylindric
 
|stem segment width=2cm;2.5cm
 
|stigma lobe coloration=white;light green
 
|style coloration=white;light green
 
|tip coloration=yellow;golden
 
|tree architecture=branched
 
|tree atypical some measurement=2m;4m
 
|tree some measurement=1m;2m
 
|tubercle prominence=prominent
 
|tubercle some measurement=1.5cm;4.5cm
 
|tubercle width=narrow;broad
 
|umbilicus depth=shallow
 
|wool coloration=gray-black;yellowish;tan
 
|wool life cycle=aging
 
 
}}<!--
 
}}<!--
  
-->[[Category:Treatment]][[Category:Cylindropuntia]]
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[[Category:Treatment]]
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[[Category:Cylindropuntia]]
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[[Category:Revised Since Print]]

Latest revision as of 18:01, 6 November 2020

Trees or shrubs, sparsely to densely branched, 1–2(–4) m. Stem segments firmly attached, cylindric, 10–30(–50) × 2–2.5(–3) cm; tubercles prominent, narrow to broad, 1.5–4.5 cm; areoles elliptic to subcircular, 4–5.5 × 4–5 mm; wool white or yellowish to tan, aging gray-black. Spines 6–20(–30+) per areole, often accompanied by 0–5 short bristlelike spines at areole margins, usually in distal areoles; major abaxial spines deflexed to divergent, subterete to flattened; major adaxial spines ascending-divergent, terete, yellow or tan to deep red-brown, aging gray, the central spine longest, 12–30(–38) mm; sheaths white to grayish with yellow to golden tips, baggy to tight fitting. Glochids in inconspicuous to small adaxial tuft, yellow to brown, 0.5–2 mm. Flowers: inner tepals bright yellow to bronze to brick red, spatulate, 20–30 mm, apiculate-emarginate; filaments red; anthers yellow; style and stigma lobes white to light green. Fruits tan at maturity, obconic to ellipsoid, 15–35 × 15–20 mm, dry, tuberculate, densely to sparsely spiny (rarely spineless), with apical flange above shallow umbilicus; basal tubercles longest; areoles 12–30. Seeds pale yellow to tan, angular or squarish in outline, warped, 3.5–5 × 3–4.5 mm, sides smooth, each with 2–4 large depressions; girdle smooth.

Distribution

V4 222-distribution-map.gif

Ariz., Calif., Nev., Utah, n Mexico.

Discussion

Varieties 4 (4 in the flora).

Cylindropuntia acanthocarpa hybridizes with C. abyssi, C. echinocarpa (= C. ×deserta), C. bigelovii (= C. ×campii), C. leptocaulis (see C. ×tetracantha), C. ×multigeniculata, C. spinosior, C. versicolor, and C. whipplei (= C. congesta). Hybrids of C. acanthocarpa and C. ×multigeniculata are more open in habit, with longer stem segments than C. ×multigeniculata and with reddish filaments like C. acanthocarpa.

Selected References

None.

Key

1 Tubercles 2.5-4.5 cm, broad; stem segments gray-green; spines of adjacent areoles not or barely interlaced, 6-11(-14) per areole; fruits bearing few spines or sometimes spineless Cylindropuntia acanthocarpa var. thornbe
1 Tubercles (1-)1.5-2.5(-3) cm, broad to narrow; stem segments green to yellow-green; spines of adjacent areoles interlaced; fruits densely spiny, sometimes burlike > 2
2 Abaxial 5-7 spines deflexed, usually subparallel, whitish to light brown; tubercles broad, oval; shrubs with few, ascending branches Cylindropuntia acanthocarpa var. acantho
2 Abaxial (14-)18-30+ spines deflexed to divergent, spreading, pale yellow to red-brown; tubercles narrowly elliptic; shrubs with few to many, spreading branches > 3
3 Branches usually at acute angles; stem segments yellowish green, usually long, commonly 15-30 cm; spines 18-30+, yellow to tan, sheaths baggy; inner tepals yellow Cylindropuntia acanthocarpa var. colorad
3 Branches usually at obtuse angles; stem segments green to dark green, usually short, commonly 8 -20 cm; spines 14-18(-23) per areole, yellow or red-brown, sheaths tight; inner tepals yellow to bronze to brick red Cylindropuntia acanthocarpa var. major