Difference between revisions of "Castanea"

Miller

Gard. Dict. Abr. ed. 4. 1754.

Common names: Chestnut châtaignier
Etymology: Classical Latin, from Greek kastanaion karuon, nut from Castania, probably referring either to Kastanaia in Pontus or Castana in Thessaly
Treatment appears in FNA Volume 3.
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--><span class="statement" id="st-d0_s0" data-properties="tree duration;tree architecture;shrub duration;shrub architecture"><b>Trees </b>or shrubs, winter-deciduous, sometimes rhizomatous.</span> <span class="statement" id="st-d0_s1" data-properties="terminal bud presence;pseudoterminal bud shape;outer scale quantity;outer scale size;outer scale arrangement;inner scale quantity;inner scale arrangement"><b>Terminal </b>buds absent, pseudoterminal bud (axillary bud of youngest leaf) ovoid, with 2 unequal opposite outer scales enclosing several imbricate inner scales.</span> <span class="statement" id="st-d0_s2" data-properties="stipule prominence;stipule duration;growth life cycle"><b>Leaves:</b> stipules prominent on new growth, soon deciduous.</span> <span class="statement" id="st-d0_s3" data-properties="leaf-blade width;leaf-blade texture;secondary-vein architecture;secondary-vein arrangement;tooth shape;awn shape;awn development"><b>Leaf-</b>blade thin, somewhat leathery, secondary-veins unbranched, ±parallel, extending to margin, each vein ending in sharp tooth or well-developed awn.</span> <span class="statement" id="st-d0_s4" data-properties="inflorescence architecture;inflorescence reproduction;inflorescence position;inflorescence architecture;inflorescence orientation;inflorescence texture;inflorescence fragility"><b>Inflorescences </b>staminate or androgynous, axillary, spicate, erect, rigid or flexible;</span> <span class="statement" id="st-d0_s5" data-properties="inflorescence reproduction;flower architecture;flower architecture">androgynous inflorescences with pistillate cupules/flowers toward base and staminate flowers distally.</span> <span class="statement" id="st-d0_s6" data-properties="flower architecture;sepal fusion"><b>Staminate </b>flowers: sepals distinct;</span> <span class="statement" id="st-d0_s7" data-properties="flower architecture;stamen atypical quantity;stamen quantity;pistillode position relational;pistillode texture;hair pubescence">stamens 12 (-18), typically surrounding indurate pistillode covered with silky hairs.</span> <span class="statement" id="st-d0_s8" data-properties="flower architecture;flower quantity"><b>Pistillate </b>flowers 1-3 per cupule;</span> <span class="statement" id="st-d0_s9" data-properties="sepal fusion">sepals distinct;</span> <span class="statement" id="st-d0_s10" data-properties="style atypical quantity;style quantity">carpels and styles typically 6 (-9).</span> <span class="statement" id="st-d0_s11" data-properties=""><b>Fruits:</b> maturation in 1st year following pollination (termed annual by many authors);</span> <span class="statement" id="st-d0_s12" data-properties="cupule architecture;valve fusion;valve life cycle;valve architecture or shape;spine architecture;spine position relational;hair architecture">cupule 2-4-valved, valves connate marginally until maturity, ±completely enclosing nut (s), spiny, spines irregularly branched, often interlocking, densely or sparsely covered in simple hairs;</span> <span class="statement" id="st-d0_s13" data-properties="nut quantity;nut shape;nut quantity;central nut size;central nut shape;central nut architecture or arrangement or growth form;central nut shape;central nut architecture">nuts 1-3 per cupule, planoconvex, or if 3, then central nut often reduced and flattened, or if solitary, then often rounded in cross-section, not winged, adjacent nuts not separated by internal cupule valves.</span> <span class="statement" id="st-d0_s14" data-properties="nut arrangement;nut arrangement;x chromosome quantity">x = 12.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Trees </b>or shrubs, winter-deciduous, sometimes rhizomatous. <b>Terminal</b> buds absent, pseudoterminal bud (axillary bud of youngest leaf) ovoid, with 2 unequal opposite outer scales enclosing several imbricate inner scales. <b>Leaves</b>: stipules prominent on new growth, soon deciduous. <b>Leaf</b> blade thin, somewhat leathery, secondary veins unbranched, ±parallel, extending to margin, each vein ending in sharp tooth or well-developed awn. <b>Inflorescences</b> staminate or androgynous, axillary, spicate, erect, rigid or flexible; androgynous inflorescences with pistillate cupules/flowers toward base and staminate flowers distally. <b>Staminate</b> flowers: sepals distinct; stamens 12(-18), typically surrounding indurate pistillode covered with silky hairs. <b>Pistillate</b> flowers 1-3 per cupule; sepals distinct; carpels and styles typically 6(-9). <b>Fruits</b>: maturation in 1st year following pollination (termed annual by many authors); cupule 2-4-valved, valves connate marginally until maturity, ±completely enclosing nut(s), spiny, spines irregularly branched, often interlocking, densely or sparsely covered in simple hairs; nuts 1-3 per cupule, plano-convex, or if 3, then central nut often reduced and flattened, or if solitary, then often rounded in cross section, not winged, adjacent nuts not separated by internal cupule valves. <b>x</b> = 12.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
|distribution=North America;Europe;Asia
+
|distribution=North America;Europe;Asia.
 
|discussion=<p>Species ca. 8-10 (3 in the flora, often interpreted as 2).</p><!--
 
|discussion=<p>Species ca. 8-10 (3 in the flora, often interpreted as 2).</p><!--
 
--><p>As evidenced by United States breeding programs, all species are probably interfertile (including American × Asian species). Local morphologic intergradation between species is to be expected.</p>
 
--><p>As evidenced by United States breeding programs, all species are probably interfertile (including American × Asian species). Local morphologic intergradation between species is to be expected.</p>
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|basionyms=
 
|basionyms=
 
|family=Fagaceae
 
|family=Fagaceae
|distribution=North America;Europe;Asia
+
|distribution=North America;Europe;Asia.
 
|reference=hardin1985a;johnson1988b;paillet1989a;paillet1993a;tucker1975a
 
|reference=hardin1985a;johnson1988b;paillet1989a;paillet1993a;tucker1975a
 
|publication title=Gard. Dict. Abr. ed.
 
|publication title=Gard. Dict. Abr. ed.
 
|publication year=1754
 
|publication year=1754
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V3/V3_684.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V3/V3_684.xml
 
|genus=Castanea
 
|genus=Castanea
|awn development=well-developed
 
|awn shape=sharp
 
|central nut architecture=not winged
 
|central nut architecture or arrangement or growth form=solitary
 
|central nut shape=rounded;flattened
 
|central nut size=reduced
 
|cupule architecture=2-4-valved
 
|flower architecture=pistillate;staminate;staminate;staminate;pistillate
 
|flower quantity=1;3
 
|growth life cycle=new
 
|hair architecture=simple
 
|hair pubescence=silky
 
|inflorescence architecture=spicate;staminate
 
|inflorescence fragility=pliable
 
|inflorescence orientation=erect
 
|inflorescence position=axillary
 
|inflorescence reproduction=androgynous;androgynous
 
|inflorescence texture=rigid
 
|inner scale arrangement=imbricate
 
|inner scale quantity=several
 
|leaf-blade texture=leathery
 
|leaf-blade width=thin
 
|nut arrangement=not separated;adjacent
 
|nut quantity=3;1;3
 
|nut shape=planoconvex
 
|outer scale arrangement=opposite
 
|outer scale quantity=2
 
|outer scale size=unequal
 
|pistillode position relational=surrounding
 
|pistillode texture=indurate
 
|pseudoterminal bud shape=ovoid
 
|secondary-vein architecture=unbranched
 
|secondary-vein arrangement=parallel
 
|sepal fusion=distinct;distinct
 
|shrub architecture=rhizomatous
 
|shrub duration=winter-deciduous
 
|spine architecture=branched
 
|spine position relational=interlocking
 
|stamen atypical quantity=12;18
 
|stamen quantity=12
 
|stipule duration=deciduous
 
|stipule prominence=prominent
 
|style atypical quantity=6;9
 
|style quantity=6
 
|terminal bud presence=absent
 
|tooth shape=sharp
 
|tree architecture=rhizomatous
 
|tree duration=winter-deciduous
 
|valve architecture or shape=spiny
 
|valve fusion=connate
 
|valve life cycle=maturity
 
|x chromosome quantity=12
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Fagaceae]]
 
-->[[Category:Treatment]][[Category:Fagaceae]]

Revision as of 14:47, 27 July 2019

Trees or shrubs, winter-deciduous, sometimes rhizomatous. Terminal buds absent, pseudoterminal bud (axillary bud of youngest leaf) ovoid, with 2 unequal opposite outer scales enclosing several imbricate inner scales. Leaves: stipules prominent on new growth, soon deciduous. Leaf blade thin, somewhat leathery, secondary veins unbranched, ±parallel, extending to margin, each vein ending in sharp tooth or well-developed awn. Inflorescences staminate or androgynous, axillary, spicate, erect, rigid or flexible; androgynous inflorescences with pistillate cupules/flowers toward base and staminate flowers distally. Staminate flowers: sepals distinct; stamens 12(-18), typically surrounding indurate pistillode covered with silky hairs. Pistillate flowers 1-3 per cupule; sepals distinct; carpels and styles typically 6(-9). Fruits: maturation in 1st year following pollination (termed annual by many authors); cupule 2-4-valved, valves connate marginally until maturity, ±completely enclosing nut(s), spiny, spines irregularly branched, often interlocking, densely or sparsely covered in simple hairs; nuts 1-3 per cupule, plano-convex, or if 3, then central nut often reduced and flattened, or if solitary, then often rounded in cross section, not winged, adjacent nuts not separated by internal cupule valves. x = 12.

Distribution

North America, Europe, Asia.

Discussion

Species ca. 8-10 (3 in the flora, often interpreted as 2).

As evidenced by United States breeding programs, all species are probably interfertile (including American × Asian species). Local morphologic intergradation between species is to be expected.

Key

1 Cupules 4-valved, enclosing 3 flowers/fruits; leaf blade abaxially without stellate trichomes, appearing glabrous, with minute multicellular glands, these often embedded on blade, and simple trichomes on veins; nut obovate, flattened at least on 1 side, beak thin, flexible, to 8 mm or more excluding styles. Castanea dentata
1 Cupules 2-valved, enclosing 1 flower/fruit; leaf blade abaxially bearing stellate trichomes (occasionally visible only with magnification), often with simple trichomes on veins; nut round in cross section, ovoid-conic, beak less than 3mm excluding styles. > 2
2 Longest spines of cupule often exceeding 10 mm; young twigs glabrous; petiole usually 8–10(–15) mm; bark brownish, moderately to deeply fissured. Castanea ozarkensis
2 Longest spines of cupule usually less than 10 mm; young twigs puberulent (sometimes glabrate with age); petiole usually 3–7(–10) mm; bark gray to brown, smooth, not fissured or only shallowly fissured. Castanea pumila
... more about "Castanea"
Kevin C. Nixon +
Miller +
Chestnut +  and châtaignier +
North America +, Europe +  and Asia. +
Classical Latin, from Greek kastanaion karuon, nut from Castania, probably referring either to Kastanaia in Pontus or Castana in Thessaly +
Gard. Dict. Abr. ed. +
hardin1985a +, johnson1988b +, paillet1989a +, paillet1993a +  and tucker1975a +
Castanea +
Fagaceae +