Difference between revisions of "Athyrium filix-femina"

(Linnaeus) Roth ex Mertens

Arch. Bot. (Leipzig) 2(1): 106. 1799.

Common names: Lady fern athyrie fougère-femelle
Basionym: Polypodium filix-femina Linnaeus
Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties="stem orientation;stem orientation"><b>Stems </b>short-creeping or ascending.</span> <span class="statement" id="st-d0_s1" data-properties="petiole coloration;petiole some measurement;base coloration;base coloration;base shape;row count"><b>Petiole </b>straw-colored distally, 7–60 cm, base dark redbrown or black, swollen, with 2 rows of teeth;</span> <span class="statement" id="st-d0_s2" data-properties="scale coloration;scale coloration;scale coloration;scale shape;scale shape;scale shape;scale length;scale width">scales light to dark-brown, linear to ovatelanceolate, 7–20 × 1–5 mm.</span> <span class="statement" id="st-d0_s3" data-properties="blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade length;blade width;blade growth form or texture;blade shape;margin pubescence or texture;apex shape"><b>Blade </b>elliptic, lanceolate to oblanceolate, 2-pinnate to 2-pinnate-pinnatifid, 18–30 × 5–50 cm, herbaceous but with cartilaginous margin, narrowed to base, apex acuminate.</span> <span class="statement" id="st-d0_s4" data-properties="pinna architecture;pinna architecture;pinna architecture;pinna shape;pinna shape;pinna shape;apex shape"><b>Pinnae </b>sessile to short-stalked, linear-oblong to lanceolate, apex acuminate.</span> <span class="statement" id="st-d0_s5" data-properties="pinnule shape;segment shape;segment shape;segment shape;margin architecture or shape"><b>Pinnules </b>pinnatifid, segments oblong-linear to narrowly deltate, margins serrate.</span> <span class="statement" id="st-d0_s6" data-properties="rachis pubescence;rachis pubescence;costa pubescence;costule pubescence"><b>Rachis,</b> costae, and costules glabrous or with glands or hairs.</span> <span class="statement" id="st-d0_s7" data-properties="vein architecture or shape"><b>Veins </b>pinnate.</span> <span class="statement" id="st-d0_s8" data-properties="sorus course;sorus shape;sorus shape"><b>Sori </b>straight, hooked at distal end, or horseshoe-shaped;</span> <span class="statement" id="st-d0_s9" data-properties="indusium architecture;indusium architecture">indusia dentate or ciliate.</span><!--
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--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>short-creeping or ascending. <b>Petiole</b> straw-colored distally, 7–60 cm, base dark red-brown or black, swollen, with 2 rows of teeth; scales light to dark brown, linear- to ovate-lanceolate, 7–20 × 1–5 mm. <b>Blade</b> elliptic, lanceolate to oblanceolate, 2-pinnate to 2-pinnate-pinnatifid, 18–30 × 5–50 cm, herbaceous but with cartilaginous margin, narrowed to base, apex acuminate. <b>Pinnae</b> sessile to short-stalked, linear-oblong to lanceolate, apex acuminate. <b>Pinnules</b> pinnatifid, segments oblong-linear to narrowly deltate, margins serrate. <b>Rachis</b>, costae, and costules glabrous or with glands or hairs. <b>Veins</b> pinnate. <b>Sori</b> straight, hooked at distal end, or horseshoe-shaped; indusia dentate or ciliate.</span><!--
  
 
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|publication year=1799
 
|publication year=1799
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_40.xml
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_40.xml
 
|genus=Athyrium
 
|genus=Athyrium
 
|species=Athyrium filix-femina
 
|species=Athyrium filix-femina
|apex shape=acuminate;acuminate
 
|base coloration=black;dark redbrown
 
|base shape=swollen
 
|blade growth form or texture=herbaceous
 
|blade length=18cm;30cm
 
|blade shape=narrowed;lanceolate;oblanceolate 2-pinnate
 
|blade width=5cm;50cm
 
|costa pubescence=glabrous
 
|costule pubescence=glabrous
 
|indusium architecture=ciliate;dentate
 
|margin architecture or shape=serrate
 
|margin pubescence or texture=cartilaginous
 
|petiole coloration=straw-colored
 
|petiole some measurement=7cm;60cm
 
|pinna architecture=sessile;short-stalked
 
|pinna shape=linear-oblong;lanceolate
 
|pinnule shape=pinnatifid
 
|rachis pubescence=with glands or hairs;glabrous
 
|row count=2
 
|scale coloration=light;dark-brown
 
|scale length=7mm;20mm
 
|scale shape=linear;ovatelanceolate
 
|scale width=1mm;5mm
 
|segment shape=oblong-linear;narrowly deltate
 
|sorus course=straight
 
|sorus shape=horseshoe-shaped;hooked
 
|stem orientation=ascending;short-creeping
 
|vein architecture or shape=pinnate
 
 
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Revision as of 14:19, 27 July 2019

Stems short-creeping or ascending. Petiole straw-colored distally, 7–60 cm, base dark red-brown or black, swollen, with 2 rows of teeth; scales light to dark brown, linear- to ovate-lanceolate, 7–20 × 1–5 mm. Blade elliptic, lanceolate to oblanceolate, 2-pinnate to 2-pinnate-pinnatifid, 18–30 × 5–50 cm, herbaceous but with cartilaginous margin, narrowed to base, apex acuminate. Pinnae sessile to short-stalked, linear-oblong to lanceolate, apex acuminate. Pinnules pinnatifid, segments oblong-linear to narrowly deltate, margins serrate. Rachis, costae, and costules glabrous or with glands or hairs. Veins pinnate. Sori straight, hooked at distal end, or horseshoe-shaped; indusia dentate or ciliate.

Distribution

North America, Mexico, Central America, South America, Europe, Asia.

Discussion

Athyrium filix-femina is circumboreal, and this or closely related species extend into Mexico, Central America, and South America. The delimitation and infraspecific classification of A. filix-femina need detailed study.

Varieties ca. 5 (4 in the flora).

Selected References

None.

Key

1 Petiole scales more than 1 cm; blades ca. 2 times length of petioles, elliptic to oblanceolate. > 2
1 Petiole scales 1 cm or less; blades 1-1.5 times length of petioles, elliptic to lanceolate. > 3
2 Pinnules narrowly deltate or oblong-lanceolate, nearly equilateral at base; indusia long-ciliate; spores yellow. Athyrium filix-femina var. cyclosorum
2 Pinnules linear-oblong or linear-lanceolate, inequilateral at base; indusia dentate or long-ciliate; spores brown. Athyrium filix-femina var. californicum
3 Petiole scales brown to dark brown; blades elliptic, narrowed to base, broadest near or just below middle; pinnae sessile or short-stalked; pinnules linear to oblong; indusia not glandular; spores yellow. Athyrium filix-femina var. angustum
3 Petiole scales light brown to brown; blades ovate-lanceolate to lanceolate, slightly narrowed to base, broadest just above base; pinnae usually stalked; pinnules oblong-lanceolate to narrowly deltate; indusia glandular or not; spores dark brown. Athyrium filix-femina var. asplenioides