Hypericum majus
Mem. Torrey Bot. Club 5: 225. 1894.
Herbs perennial, erect, branching at base and in inflorescence, branches relatively few, 0.5–7 dm. Stems: internodes 4-angled. Leaves spreading, sessile or distal subamplexicaul; blade lanceolate to narrowly oblong-elliptic or (proximal) oblanceolate, 10–45 × (2–)6–12 mm, papery to membranous, margins plane, apex acute to rounded, basal or near-basal veins (3–)5–7, midrib with to 4 pairs of branches. Inflorescences corymbiform to cylindric, 3–30-flowered, usually compact, branching mostly dichasial. Flowers 6–7 mm diam.; sepals lanceolate to narrowly elliptic, equal, 3.5–6.5 × 0.8–1.5 mm, margins sometimes ciliate, not setulose-ciliate, apex acute; petals golden yellow, sometimes red-veined, oblanceolate, 3.5–6 mm; stamens 12–21, obscurely 5-fascicled; styles 0.6–1 mm; stigmas broadly capitate. Capsules narrowly conic-ellipsoid, 4–8 × 2.5–3.5 mm, broadest proximal to middle. Seeds 0.5–0.7 mm; testa finely linear-scalariform. 2n = 16.
Phenology: Flowering summer (Jun–Sep).
Habitat: Fens, marshes, ditches, lake and stream margins, other damp habitats
Elevation: 0–1200 m
Distribution
Alta., B.C., Man., N.B., N.S., Ont., P.E.I., Que., Sask., Colo., Conn., Idaho, Ill., Ind., Iowa, Kans., Maine, Mass., Mich., Minn., Mo., Mont., Nebr., N.H., N.J., N.Y., N.Dak., Ohio, Oreg., Pa., R.I., S.Dak., Vt., Wash., Wis., introduced in Europe (France, Germany), e Asia (Japan).
Discussion
Hypericum majus was the western member of a vicariant species pair, differing from the originally eastern member (H. canadense) by the broader leaves, usually more-congested inflorescence, and larger flowers. These species became sympatric in glaciated northeastern North America and now hybridize freely, notably in Wisconsin (F. H. Utech and H. H. Iltis 1970). Hybrids are intermediate in form between the parents and have also been recorded from Connecticut, Massachusetts, and New Hampshire. Hypericum majus hybridizes also with H. mutilum, with subsp. mutilum in Maine, and with subsp. boreale in Michigan and Wisconsin.
Selected References
None.