Erythranthe grandis
Phytoneuron 2012-39: 43. 2012.
Perennials, rhizomatous, sometimes rooting at proximal nodes. Stems erect, sometimes decumbent basally, branched, often fistulose, (25–)50–120(–160) cm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs. Leaves basal and cauline, basal usually not persistent, bracteate in inflorescence; petiole 10–80 mm, gradually reduced distally; blade subpinnately, sometimes palmately, 5–7-veined, ovate to broadly elliptic, 25–60 × 20–40(–60) mm, usually 1–2 times longer than wide, base truncate or truncate-cuneate to subcordate, margins crenulate to dentate, proximally sometimes sublyrate, apex rounded to obtuse, surfaces of distals densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs. Flowers herkogamous, 8–26, mostly from distal nodes, usually in bracteate racemes. Fruiting pedicels 10–35 mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs. Fruiting calyces straight-erect or nodding 45–100º, ovate-campanulate, inflated, sagittally compressed, 15–22(–25) mm, densely hirsutulous to softly hirtellous-puberulent to pilose-hirsutulous, hairs usually crinkly, and eglandular or with a mixture of hirtellous-puberulent and stipitate-glandular hairs, sometimes ± stipitate-glandular or glandular-villous without hirtellous-puberulent hairs, throat closing. Corollas yellow, red-dotted within, bilaterally symmetric, bilabiate; tube-throat broadly funnelform, (14–)16–24 mm, exserted (8–)10–15 mm beyond calyx margin; limb broadly expanded. Styles hirtellous. Anthers included, glabrous. Capsules included, 8–12 mm. 2n = 28.
Phenology: Flowering (Apr–)May–Jul(–Sep).
Habitat: Beaches, dunes, coastal bluffs, wet cliff faces, mud flats and seeps, marshes, drainage ditches, creeks, rarely in coastal sage scrub.
Elevation: 0–200(–800) m.
Discussion
The densely, evenly puberulent vestiture of pedicels, calyces, and distal stems usually is diagnostic, especially in combination with the large flowers (corollas and mature calyces) and tall stature. Plants from scattered collections are much shorter than normal but have large corollas and characteristic vestiture.
Erythranthe grandis characteristically occurs in coastal localities from southern California to northern Oregon but also is found in inland localities and habitats near the coast but well away from salt spray.
Selected References
None.