Difference between revisions of "Sclerocactus cloverae"

K. D. Heil & J. M. Porter

Haseltonia 2: 31. 1994.

Common names: Clover eagle-claw cactus Clover’s fishhook cactus
Basionym: Sclerocactus whipplei var. heilii CastetterSclerocactus whipplei var. reevesii Castetter
Synonyms: Pediocactus cloverae (K. D. Heil & J. M. Porter) HaldaSclerocactus cloverae subsp. brackii K. D. Heil & J. M. Porter
Treatment appears in FNA Volume 4. Treatment on page 202. Mentioned on page 199, 201.
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--><span class="statement" id="st-d0_s0" data-properties="stem architecture;stem coloration;stem shape;stem shape;stem shape;stem atypical length;stem length;stem atypical width;stem width"><b>Stems </b>unbranched (occasionally 2–3-branched near base), green, ovoid to elongate-cylindric, 2.9–25 (–35) × 2.8–12.5 (–20) cm;</span> <span class="statement" id="st-d0_s1" data-properties="rib atypical count;rib atypical count;rib count;rib development;tubercle prominence">ribs usually (11–) 13 (–15), well developed, tubercles evident on ribs.</span> <span class="statement" id="st-d0_s2" data-properties=""><b>Spines </b>obscuring stems;</span> <span class="statement" id="st-d0_s3" data-properties="spine arrangement;spine count;spine shape;spine shape;spine shape;spine length;spine width">radial spines 4–6 per areole, acicular, elliptic or rhombic in cross-section, 19 × 1.3–2 mm;</span> <span class="statement" id="st-d0_s4" data-properties="central spine count;areole count">central spines 6–9 per areole, usually 8;</span> <span class="statement" id="st-d0_s5" data-properties="abaxial central spine count;abaxial central spine orientation;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine shape;abaxial central spine shape;abaxial central spine shape;abaxial central spine atypical length;abaxial central spine length;abaxial central spine width">abaxial central spine usually 1 per areole, porrect, straw colored to brown, highlighted with purple or red, terete or somewhat angled, hooked, (15–) 30–46 × 1.5 mm;</span> <span class="statement" id="st-d0_s6" data-properties="lateral central spine count;lateral central spine shape;abaxial spine height or length or size">lateral central spines 5–8 per areole, similar to abaxial but slightly shorter and usually not hooked;</span> <span class="statement" id="st-d0_s7" data-properties="adaxial central spine orientation;adaxial central spine coloration;adaxial central spine coloration;adaxial central spine course;adaxial central spine course;adaxial central spine shape;adaxial central spine prominence;adaxial central spine shape;adaxial central spine length;adaxial central spine width">adaxial central erect, white or straw colored, straight or curved, angled to flat, somewhat inconspicuous, triangular in cross-section, 25–55 × 1–2 mm.</span> <span class="statement" id="st-d0_s8" data-properties="flower shape;flower shape;flower shape;flower atypical length;flower length;flower atypical width;flower width"><b>Flowers </b>narrowly funnelform to campanulate, 2.5–3.5 (–4) × 1.6–3.1 (–3.6) cm;</span> <span class="statement" id="st-d0_s9" data-properties="outer tepal coloration;outer tepal coloration;outer tepal coloration;outer tepal coloration;outer tepal coloration;outer tepal coloration;outer tepal coloration;outer tepal coloration;outer tepal size;outer tepal shape;outer tepal length;outer tepal width;midstripe coloration;margin texture;margin shape;margin shape;apex shape">outer tepals with greenish to purple with brownish midstripes, pink, purple, or whitish margins, larger ones oblanceolate, 10–18 × 4–7.5 mm, margins membranous and crisped or minutely toothed, apex mucronate;</span> <span class="statement" id="st-d0_s10" data-properties="inner tepal coloration or density;inner tepal coloration;largest tepal shape;largest tepal atypical length;largest tepal length;largest tepal width;margin shape;apex shape">inner tepals purple, sometimes suffused with brown, largest tepals oblanceolate, 15–22 (–30) × 4–6 mm, margins irregularly toothed, apex mucronate;</span> <span class="statement" id="st-d0_s11" data-properties="filament coloration;filament coloration;filament coloration;filament coloration;filament coloration">filaments white, tinged with pink to pink-purple;</span> <span class="statement" id="st-d0_s12" data-properties="anther coloration">anthers yellow;</span> <span class="statement" id="st-d0_s13" data-properties="ovary relief;ovary architecture or pubescence or relief">ovary minutely papillate, appearing smooth.</span> <span class="statement" id="st-d0_s14" data-properties="fruit dehiscence;fruit dehiscence;fruit coloration;fruit coloration;fruit coloration;fruit coloration;fruit length;fruit width;fruit condition or texture;pore position;pore life cycle"><b>Fruits </b>irregularly dehiscent or dehiscent through basal abscission pore, green to tan, sometimes suffused with pink, 7–15 × 5–12 mm, dry;</span> <span class="statement" id="st-d0_s15" data-properties="scale count;scale texture;scale architecture;scale shape;scale shape">scales few, membranous, scarious-margined, minutely toothed or fringed.</span> <span class="statement" id="st-d0_s16" data-properties="seed coloration;seed coloration;seed length;seed width"><b>Seeds </b>brown or black, 1.2–2.5 × 1.9–3.5 mm;</span> <span class="statement" id="st-d0_s17" data-properties="papilla shape">testa with rounded papillae.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>unbranched (occasionally 2–3-branched near base), green, ovoid to elongate-cylindric, 2.9–25(–35) × 2.8–12.5(–20) cm; ribs usually (11–)13(–15), well developed, tubercles evident on ribs. <b>Spines</b> obscuring stems; radial spines 4–6 per areole, acicular, elliptic or rhombic in cross section, 19 × 1.3–2 mm; central spines 6–9 per areole, usually 8; abaxial central spine usually 1 per areole, porrect, straw colored to brown, highlighted with purple or red, terete or somewhat angled, hooked, (15–)30–46 × 1.5 mm; lateral central spines 5–8 per areole, similar to abaxial but slightly shorter and usually not hooked; adaxial central erect, white or straw colored, straight or curved, angled to flat, somewhat inconspicuous, triangular in cross section, 25–55 × 1–2 mm. <b>Flowers</b> narrowly funnelform to campanulate, 2.5–3.5(–4) × 1.6–3.1(–3.6) cm; outer tepals with greenish to purple with brownish midstripes, pink, purple, or whitish margins, larger ones oblanceolate, 10–18 × 4–7.5 mm, margins membranous and crisped or minutely toothed, apex mucronate; inner tepals purple, sometimes suffused with brown, largest tepals oblanceolate, 15–22(–30) × 4–6 mm, margins irregularly toothed, apex mucronate; filaments white, tinged with pink to pink-purple; anthers yellow; ovary minutely papillate, appearing smooth. <b>Fruits</b> irregularly dehiscent or dehiscent through basal abscission pore, green to tan, sometimes suffused with pink, 7–15 × 5–12 mm, dry; scales few, membranous, scarious-margined, minutely toothed or fringed. <b>Seeds</b> brown or black, 1.2–2.5 × 1.9–3.5 mm; testa with rounded papillae.</span><!--
  
 
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|phenology=Flowering late Apr-early Jun.
 
|habitat=Sandy, gravelly, or clay hills, mesas, and washes, desert grasslands, saltbush, sagebrush, and rabbitbrush flats, pinyon-juniper woodlands
 
|habitat=Sandy, gravelly, or clay hills, mesas, and washes, desert grasslands, saltbush, sagebrush, and rabbitbrush flats, pinyon-juniper woodlands
 
|elevation=1500-2200 m
 
|elevation=1500-2200 m
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|basionyms=Sclerocactus whipplei var. heilii;Sclerocactus whipplei var. reevesii
 
|basionyms=Sclerocactus whipplei var. heilii;Sclerocactus whipplei var. reevesii
 
|family=Cactaceae
 
|family=Cactaceae
 +
|phenology=Flowering late Apr-early Jun.
 
|habitat=Sandy, gravelly, or clay hills, mesas, and washes, desert grasslands, saltbush, sagebrush, and rabbitbrush flats, pinyon-juniper woodlands
 
|habitat=Sandy, gravelly, or clay hills, mesas, and washes, desert grasslands, saltbush, sagebrush, and rabbitbrush flats, pinyon-juniper woodlands
 
|elevation=1500-2200 m
 
|elevation=1500-2200 m
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|publication year=1994
 
|publication year=1994
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_378.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V4/V4_378.xml
 
|subfamily=Cactaceae subfam. Cactoideae
 
|subfamily=Cactaceae subfam. Cactoideae
 
|genus=Sclerocactus
 
|genus=Sclerocactus
 
|species=Sclerocactus cloverae
 
|species=Sclerocactus cloverae
|abaxial central spine atypical length=15mm;30mm
 
|abaxial central spine coloration=,;highlighted with purple or highlighted with red;straw colored;brown
 
|abaxial central spine count=1
 
|abaxial central spine length=30mm;46mm
 
|abaxial central spine orientation=porrect
 
|abaxial central spine shape=hooked;angled;terete
 
|abaxial central spine width=1.5
 
|abaxial spine height or length or size=shorter
 
|adaxial central spine coloration=straw colored;white
 
|adaxial central spine course=curved;straight
 
|adaxial central spine length=25mm;55mm
 
|adaxial central spine orientation=erect
 
|adaxial central spine prominence=inconspicuous
 
|adaxial central spine shape=triangular;angled to flat
 
|adaxial central spine width=1mm;2mm
 
|anther coloration=yellow
 
|apex shape=mucronate;mucronate
 
|areole count=8
 
|central spine count=6;9
 
|filament coloration=tinged with pink;pink-purple
 
|flower atypical length=3.5cm;4cm
 
|flower atypical width=3.1cm;3.6cm
 
|flower length=2.5cm;3.5cm
 
|flower shape=narrowly funnelform;campanulate
 
|flower width=1.6cm;3.1cm
 
|fruit coloration=suffused with pink;green;tan
 
|fruit condition or texture=dry
 
|fruit dehiscence=dehiscent;dehiscent
 
|fruit length=7mm;15mm
 
|fruit width=5mm;12mm
 
|inner tepal coloration=suffused with brown
 
|inner tepal coloration or density=purple
 
|largest tepal atypical length=22mm;30mm
 
|largest tepal length=15mm;22mm
 
|largest tepal shape=oblanceolate
 
|largest tepal width=4mm;6mm
 
|lateral central spine count=5;8
 
|lateral central spine shape=hooked
 
|margin shape=toothed;toothed;crisped
 
|margin texture=membranous
 
|midstripe coloration=brownish
 
|outer tepal coloration=whitish;purple;whitish;purple;pink;greenish;purple
 
|outer tepal length=10mm;18mm
 
|outer tepal shape=oblanceolate
 
|outer tepal size=larger
 
|outer tepal width=4mm;7.5mm
 
|ovary architecture or pubescence or relief=smooth
 
|ovary relief=papillate
 
|papilla shape=rounded
 
|pore life cycle=abscission
 
|pore position=basal
 
|rib atypical count=13;15
 
|rib count=13
 
|rib development=developed
 
|scale architecture=scarious-margined
 
|scale count=few
 
|scale shape=fringed;toothed
 
|scale texture=membranous
 
|seed coloration=black;brown
 
|seed length=1.2mm;2.5mm
 
|seed width=1.9mm;3.5mm
 
|spine arrangement=radial
 
|spine count=4;6
 
|spine length=19
 
|spine shape=rhombic;elliptic;acicular
 
|spine width=1.3mm;2mm
 
|stem architecture=unbranched
 
|stem atypical length=25cm;35cm
 
|stem atypical width=12.5cm;20cm
 
|stem coloration=green
 
|stem length=2.9cm;25cm
 
|stem shape=ovoid;elongate-cylindric
 
|stem width=2.8cm;12.5cm
 
|tubercle prominence=evident
 
 
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Revision as of 14:08, 27 July 2019

Stems unbranched (occasionally 2–3-branched near base), green, ovoid to elongate-cylindric, 2.9–25(–35) × 2.8–12.5(–20) cm; ribs usually (11–)13(–15), well developed, tubercles evident on ribs. Spines obscuring stems; radial spines 4–6 per areole, acicular, elliptic or rhombic in cross section, 19 × 1.3–2 mm; central spines 6–9 per areole, usually 8; abaxial central spine usually 1 per areole, porrect, straw colored to brown, highlighted with purple or red, terete or somewhat angled, hooked, (15–)30–46 × 1.5 mm; lateral central spines 5–8 per areole, similar to abaxial but slightly shorter and usually not hooked; adaxial central erect, white or straw colored, straight or curved, angled to flat, somewhat inconspicuous, triangular in cross section, 25–55 × 1–2 mm. Flowers narrowly funnelform to campanulate, 2.5–3.5(–4) × 1.6–3.1(–3.6) cm; outer tepals with greenish to purple with brownish midstripes, pink, purple, or whitish margins, larger ones oblanceolate, 10–18 × 4–7.5 mm, margins membranous and crisped or minutely toothed, apex mucronate; inner tepals purple, sometimes suffused with brown, largest tepals oblanceolate, 15–22(–30) × 4–6 mm, margins irregularly toothed, apex mucronate; filaments white, tinged with pink to pink-purple; anthers yellow; ovary minutely papillate, appearing smooth. Fruits irregularly dehiscent or dehiscent through basal abscission pore, green to tan, sometimes suffused with pink, 7–15 × 5–12 mm, dry; scales few, membranous, scarious-margined, minutely toothed or fringed. Seeds brown or black, 1.2–2.5 × 1.9–3.5 mm; testa with rounded papillae.


Phenology: Flowering late Apr-early Jun.
Habitat: Sandy, gravelly, or clay hills, mesas, and washes, desert grasslands, saltbush, sagebrush, and rabbitbrush flats, pinyon-juniper woodlands
Elevation: 1500-2200 m

Discussion

Sclerocactus cloverae is characterized by its very dense spines and small (relative to S. parviflorus) purple flowers. Phylogenetic analyses of chloroplast DNA sequences support a close relationship among S. cloverae, S. whipplei, and S. parviflorus (J. M. Porter et al. 2000). Populations with all reproductive individuals maintaining juvenile morphology have been segregated as S. cloverae subsp. brackii K. D. Heil & J. M. Porter.

Selected References

None.

Lower Taxa

None.
... more about "Sclerocactus cloverae"
Kenneth D. Heil +  and J. Mark Porter +
K. D. Heil & J. M. Porter +
Sclerocactus whipplei var. heilii +  and Sclerocactus whipplei var. reevesii +
Clover eagle-claw cactus +  and Clover’s fishhook cactus +
Colo. +  and N.Mex. +
1500-2200 m +
Sandy, gravelly, or clay hills, mesas, and washes, desert grasslands, saltbush, sagebrush, and rabbitbrush flats, pinyon-juniper woodlands +
Flowering late Apr-early Jun. +
Pediocactus cloverae +  and Sclerocactus cloverae subsp. brackii +
Sclerocactus cloverae +
Sclerocactus +
species +