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| − | --><span class="statement" id="st-d0_s0" data-properties="shrub orientation;shrub some measurement;stem segment count"><b>Shrubs,</b> forming mats, widely spreading, 5–15 cm, in series of usually 1–2 stem segments.</span> <span class="statement" id="st-d0_s1" data-properties="root density"><b>Roots </b>diffuse.</span> <span class="statement" id="st-d0_s2" data-properties="stem segment shape;stem segment shape;stem segment atypical length;stem segment length;stem segment width"><b>Stem </b>segments short-clavate, strongly narrowed at base, 2.5–5 (–7.5) × 1.5–3 cm;</span> <span class="statement" id="st-d0_s3" data-properties="tubercle prominence;tubercle shape;tubercle atypical some measurement;tubercle some measurement;tubercle size or width;tubercle l w ratio;tubercle prominence;spine arrangement">tubercles prominent, oval, 5–10 (–16) mm, narrow, 4–6 times longer than wide, obscured by interlacing spines;</span> <span class="statement" id="st-d0_s4" data-properties="areole diam">areoles 3 mm in diam.;</span> <span class="statement" id="st-d0_s5" data-properties="wool coloration;wool coloration;wool coloration">wool white to gray.</span> <span class="statement" id="st-d0_s6" data-properties=""><b>Spines </b>7–15 per areole, primarily in distal areoles;</span> <span class="statement" id="st-d0_s7" data-properties="spine count;spine size;abaxial spine count;abaxial spine orientation;abaxial spine coloration;abaxial spine shape">major 3–5 abaxial spines deflexed, white, flattened, longest central spine daggerlike, broadly tapered, 12–35 × 1.5+ mm basally;</span> <span class="statement" id="st-d0_s8" data-properties="longest central spine shape;longest central spine shape;longest central spine length;longest central spine width;longest central spine size;adaxial spine count;adaxial spine orientation;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine coloration;adaxial spine shape;adaxial spine shape;adaxial spine shape">major 1–3 adaxial spines ascending, white, yellow to tan, angular-flattened to subterete.</span> <span class="statement" id="st-d0_s9" data-properties="glochid coloration;glochid some measurement"><b>Glochids </b>in adaxial 1/4 of areole, yellowish white, ± 4 mm.</span> <span class="statement" id="st-d0_s10" data-properties="inner tepal coloration;inner tepal some measurement"><b>Flowers:</b> inner tepals bright-yellow, to 25 mm;</span> <span class="statement" id="st-d0_s11" data-properties="filament coloration;filament coloration;filament coloration">filaments light yellow-green to white;</span> <span class="statement" id="st-d0_s12" data-properties="style coloration;stigma lobe coloration">style and stigma lobes white.</span> <span class="statement" id="st-d0_s13" data-properties="fruit coloration;fruit shape;fruit shape;fruit shape;fruit length;fruit width;fruit texture;fruit architecture;fruit pubescence"><b>Fruits </b>yellow, barrel-shaped to ellipsoid, 30–45 × 15–25 mm, fleshy, spineless, densely ± yellow-glochidiate;</span> <span class="statement" id="st-d0_s14" data-properties="areole count">areoles 35–55.</span> <span class="statement" id="st-d0_s15" data-properties=""><b>Seeds </b>yellowish white, 4.5 × 4 mm, smooth.</span> <span class="statement" id="st-d0_s16" data-properties="seed coloration;seed length;seed width;seed architecture or pubescence or relief;2n chromosome count">2n = 22.</span><!-- | + | --><span class="statement" id="st-undefined" data-properties=""><b>Shrubs,</b> forming mats, widely spreading, 5–15 cm, in series of usually 1–2 stem segments. <b>Roots</b> diffuse. <b>Stem</b> segments short-clavate, strongly narrowed at base, 2.5–5(–7.5) × 1.5–3 cm; tubercles prominent, oval, 5–10(–16) mm, narrow, 4–6 times longer than wide, obscured by interlacing spines; areoles 3 mm in diam.; wool white to gray. <b>Spines</b> 7–15 per areole, primarily in distal areoles; major 3–5 abaxial spines deflexed, white, flattened, longest central spine daggerlike, broadly tapered, 12–35 × 1.5+ mm basally; major 1–3 adaxial spines ascending, white, yellow to tan, angular-flattened to subterete. <b>Glochids</b> in adaxial 1/4 of areole, yellowish white, ± 4 mm. <b>Flowers</b>: inner tepals bright yellow, to 25 mm; filaments light yellow-green to white; style and stigma lobes white. <b>Fruits</b> yellow, barrel-shaped to ellipsoid, 30–45 × 15–25 mm, fleshy, spineless, densely ± yellow-glochidiate; areoles 35–55. <b>Seeds</b> yellowish white, 4.5 × 4 mm, smooth. <b>2n</b> = 22.</span><!-- |
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| | + | |phenology=Flowering late spring (May–Jun). |
| | |habitat=Great Plains grasslands, open pinyon-juniper woodlands, rocky or sandy soils | | |habitat=Great Plains grasslands, open pinyon-juniper woodlands, rocky or sandy soils |
| | |elevation=1800-2500 m | | |elevation=1800-2500 m |
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| | |basionyms=Opuntia clavata | | |basionyms=Opuntia clavata |
| | |family=Cactaceae | | |family=Cactaceae |
| | + | |phenology=Flowering late spring (May–Jun). |
| | |habitat=Great Plains grasslands, open pinyon-juniper woodlands, rocky or sandy soils | | |habitat=Great Plains grasslands, open pinyon-juniper woodlands, rocky or sandy soils |
| | |elevation=1800-2500 m | | |elevation=1800-2500 m |
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| | |publication year=1973 | | |publication year=1973 |
| | |special status= | | |special status= |
| − | |source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_235.xml | + | |source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V4/V4_235.xml |
| | |subfamily=Cactaceae subfam. Opuntioideae | | |subfamily=Cactaceae subfam. Opuntioideae |
| | |genus=Grusonia | | |genus=Grusonia |
| | |species=Grusonia clavata | | |species=Grusonia clavata |
| − | |2n chromosome count=22
| |
| − | |abaxial spine coloration=white
| |
| − | |abaxial spine count=3;5
| |
| − | |abaxial spine orientation=deflexed
| |
| − | |abaxial spine shape=flattened
| |
| − | |adaxial spine coloration=yellow;tan
| |
| − | |adaxial spine count=1;3
| |
| − | |adaxial spine orientation=ascending
| |
| − | |adaxial spine shape=angular-flattened;subterete
| |
| − | |areole count=35;55
| |
| − | |areole diam=3
| |
| − | |filament coloration=light yellow-green;white
| |
| − | |fruit architecture=spineless
| |
| − | |fruit coloration=yellow
| |
| − | |fruit length=30mm;45mm
| |
| − | |fruit pubescence=yellow-glochidiate
| |
| − | |fruit shape=barrel-shaped;ellipsoid
| |
| − | |fruit texture=fleshy
| |
| − | |fruit width=15mm;25mm
| |
| − | |glochid coloration=yellowish white
| |
| − | |glochid some measurement=4
| |
| − | |inner tepal coloration=bright-yellow
| |
| − | |inner tepal some measurement=0mm;25mm
| |
| − | |longest central spine length=12mm;35mm
| |
| − | |longest central spine shape=tapered;daggerlike
| |
| − | |longest central spine size=major
| |
| − | |root density=diffuse
| |
| − | |seed architecture or pubescence or relief=smooth
| |
| − | |seed coloration=yellowish white
| |
| − | |seed length=4.5
| |
| − | |seed width=4
| |
| − | |shrub orientation=spreading
| |
| − | |shrub some measurement=5cm;15cm
| |
| − | |spine arrangement=interlacing
| |
| − | |spine count=7;15
| |
| − | |spine size=major
| |
| − | |stem segment atypical length=5cm;7.5cm
| |
| − | |stem segment count=1;2
| |
| − | |stem segment length=2.5cm;5cm
| |
| − | |stem segment shape=narrowed;short-clavate
| |
| − | |stem segment width=1.5cm;3cm
| |
| − | |stigma lobe coloration=white
| |
| − | |style coloration=white
| |
| − | |tubercle atypical some measurement=10mm;16mm
| |
| − | |tubercle l w ratio=4-6
| |
| − | |tubercle prominence=obscured;prominent
| |
| − | |tubercle shape=oval
| |
| − | |tubercle size or width=narrow
| |
| − | |tubercle some measurement=5mm;10mm
| |
| − | |wool coloration=white;gray
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| | -->[[Category:Treatment]][[Category:Grusonia]] | | -->[[Category:Treatment]][[Category:Grusonia]] |
Shrubs, forming mats, widely spreading, 5–15 cm, in series of usually 1–2 stem segments. Roots diffuse. Stem segments short-clavate, strongly narrowed at base, 2.5–5(–7.5) × 1.5–3 cm; tubercles prominent, oval, 5–10(–16) mm, narrow, 4–6 times longer than wide, obscured by interlacing spines; areoles 3 mm in diam.; wool white to gray. Spines 7–15 per areole, primarily in distal areoles; major 3–5 abaxial spines deflexed, white, flattened, longest central spine daggerlike, broadly tapered, 12–35 × 1.5+ mm basally; major 1–3 adaxial spines ascending, white, yellow to tan, angular-flattened to subterete. Glochids in adaxial 1/4 of areole, yellowish white, ± 4 mm. Flowers: inner tepals bright yellow, to 25 mm; filaments light yellow-green to white; style and stigma lobes white. Fruits yellow, barrel-shaped to ellipsoid, 30–45 × 15–25 mm, fleshy, spineless, densely ± yellow-glochidiate; areoles 35–55. Seeds yellowish white, 4.5 × 4 mm, smooth. 2n = 22.
Phenology: Flowering late spring (May–Jun).
Habitat: Great Plains grasslands, open pinyon-juniper woodlands, rocky or sandy soils
Elevation: 1800-2500 m
Discussion
L. D. Benson (1982) mapped “Opuntia clavata” in northeast Arizona, but no supporting specimens have been found. The report may be based on low-growing specimens of Cylindropuntia whipplei, a cholla common in that area.
Selected References
None.
