Difference between revisions of "Cylindropuntia fulgida"

(Engelmann) F. M. Knuth in C. Backeberg and F. M. Knuth

in C. Backeberg and F. M. Knuth, Kaktus-ABC, 126. 1935.

Common names: Chain-fruit cholla
Basionym: Opuntia fulgida Engelmann
Treatment appears in FNA Volume 4. Mentioned on page 92, 101, 102, 105, 1.
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--><span class="statement" id="st-d0_s0" data-properties="tree some measurement"><b>Trees </b>1–3 m;</span> <span class="statement" id="st-d0_s1" data-properties="trunk architecture">trunk divaricately branching;</span> <span class="statement" id="st-d0_s2" data-properties="crown count;crown architecture;crown orientation">crown many branched, spreading.</span> <span class="statement" id="st-d0_s3" data-properties="stem segment arrangement;stem segment arrangement;stem segment coloration;stem segment condition;stem segment coloration;stem segment architecture or shape;stem segment position or structure subtype;stem segment fragility;stem segment atypical length;stem segment length;stem segment width"><b>Stem </b>segments whorled or subwhorled, gray-green, often drying blackish, ± spiny throughout, terminal ones easily dislodged, 6–16 (–23) × 2–3.5 cm;</span> <span class="statement" id="st-d0_s4" data-properties="tubercle orientation;tubercle shape;tubercle atypical some measurement;tubercle some measurement">tubercles salient, broadly oval, 0.8–1.3 (–1.9) cm;</span> <span class="statement" id="st-d0_s5" data-properties="areole shape;areole atypical length;areole length;areole width">areoles obdeltate, 5–7 (–10) × 2.5–4 mm;</span> <span class="statement" id="st-d0_s6" data-properties="wool coloration;wool coloration;wool coloration;wool coloration;wool coloration;wool coloration;wool coloration">wool gold to tan, aging gray to black.</span> <span class="statement" id="st-d0_s7" data-properties="spine atypical count;spine count;interlaced count;interlaced coloration;interlaced coloration;interlaced life cycle;interlaced coloration;interlaced count;interlaced coloration;interlaced coloration;interlaced life cycle;interlaced coloration;areole arrangement"><b>Spines </b>0–12 (–18) per areole, at most areoles to nearly absent, yellowish, sometimes also pale pinkish, aging brown, interlaced or not with spines of adjacent areoles;</span> <span class="statement" id="st-d0_s8" data-properties="abaxial spine orientation;abaxial spine orientation;abaxial spine orientation;abaxial spine orientation;abaxial spine shape;abaxial spine length;abaxial spine some measurement">abaxial spines erect to deflexed, spreading, flattened basally, the longest to 3.5 cm;</span> <span class="statement" id="st-d0_s9" data-properties="adaxial spine orientation;adaxial spine orientation;adaxial spine shape;adaxial spine shape;adaxial spine shape;adaxial spine length;adaxial spine some measurement">adaxial spines erect or spreading, terete to subterete, longest to 2.5 cm;</span> <span class="statement" id="st-d0_s10" data-properties="sheath coloration;sheath coloration;sheath coloration;sheath coloration">sheaths uniformly whitish, yellowish to golden, baggy.</span> <span class="statement" id="st-d0_s11" data-properties="glochid arrangement;glochid coloration;glochid some measurement"><b>Glochids </b>in adaxial tuft, sometimes also scattered along areole margins, yellow, 1–3 mm.</span> <span class="statement" id="st-d0_s12" data-properties="inner tepal orientation;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal shape;inner tepal architecture;inner tepal some measurement;inner tepal architecture or shape;inner tepal architecture or shape"><b>Flowers:</b> inner tepals usually reflexed, pink to magenta, obovate to ligulate, 12–16 mm, apiculate emarginate;</span> <span class="statement" id="st-d0_s13" data-properties="filament coloration;filament coloration;filament coloration">filaments pale-pink to magenta;</span> <span class="statement" id="st-d0_s14" data-properties="anther coloration;anther coloration;anther coloration">anthers white to cream;</span> <span class="statement" id="st-d0_s15" data-properties="style coloration">style pinkish;</span> <span class="statement" id="st-d0_s16" data-properties="stigma lobe coloration;stigma lobe coloration;stigma lobe coloration">stigma lobes whitish to pale-yellow.</span> <span class="statement" id="st-d0_s17" data-properties="fruit life cycle;fruit coloration;fruit architecture;fruit shape;fruit texture;fruit relief;fruit architecture;chain length or size;chain architecture;chain orientation"><b>Fruits </b>proliferating, forming long, branching, pendent chains, at maturity gray-green, often stipitate, obconic, fleshy, shallowly tuberculate, usually spineless;</span> <span class="statement" id="st-d0_s18" data-properties="basal fruit length;basal fruit width">basal fruits 32–55 × 23–45 mm;</span> <span class="statement" id="st-d0_s19" data-properties="terminal fruit length;terminal fruit width">terminal fruits 2–3.3 × 1.3–2.3 cm;</span> <span class="statement" id="st-d0_s20" data-properties="tubercle prominence">tubercles becoming obscure;</span> <span class="statement" id="st-d0_s21" data-properties="umbilicus some measurement;umbilicus depth">umbilicus to 8 mm deep;</span> <span class="statement" id="st-d0_s22" data-properties="areole count">areoles 18–35.</span> <span class="statement" id="st-d0_s23" data-properties="seed coloration;seed coloration;seed coloration;seed arrangement or shape;seed architecture or course;seed shape;seed length;seed width;depression count;depression size;hilum shape"><b>Seeds </b>pale-yellow to brownish, angular to very irregular in outline, warped, 1.9 × 1.5–3.5 mm, sides with 1–2 large depressions, hilum pointed;</span> <span class="statement" id="st-d0_s24" data-properties="girdle architecture or pubescence or relief">girdle smooth.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Trees </b>1–3 m; trunk divaricately branching; crown many branched, spreading. <b>Stem</b> segments whorled or subwhorled, gray-green, often drying blackish, ± spiny throughout, terminal ones easily dislodged, 6–16(–23) × 2–3.5 cm; tubercles salient, broadly oval, 0.8–1.3(–1.9) cm; areoles obdeltate, 5–7(–10) × 2.5–4 mm; wool gold to tan, aging gray to black. <b>Spines</b> 0–12(–18) per areole, at most areoles to nearly absent, yellowish, sometimes also pale pinkish, aging brown, interlaced or not with spines of adjacent areoles; abaxial spines erect to deflexed, spreading, flattened basally, the longest to 3.5 cm; adaxial spines erect or spreading, terete to subterete, longest to 2.5 cm; sheaths uniformly whitish, yellowish to golden, baggy. <b>Glochids</b> in adaxial tuft, sometimes also scattered along areole margins, yellow, 1–3 mm. <b>Flowers</b>: inner tepals usually reflexed, pink to magenta, obovate to ligulate, 12–16 mm, apiculate emarginate; filaments pale pink to magenta; anthers white to cream; style pinkish; stigma lobes whitish to pale yellow. <b>Fruits</b> proliferating, forming long, branching, pendent chains, at maturity gray-green, often stipitate, obconic, fleshy, shallowly tuberculate, usually spineless; basal fruits 32–55 × 23–45 mm; terminal fruits 2–3.3 × 1.3–2.3 cm; tubercles becoming obscure; umbilicus to 8 mm deep; areoles 18–35. <b>Seeds</b> pale yellow to brownish, angular to very irregular in outline, warped, 1.9 × 1.5–3.5 mm, sides with 1–2 large depressions, hilum pointed; girdle smooth.</span><!--
  
 
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|publication year=1935
 
|publication year=1935
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_200.xml
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V4/V4_200.xml
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|genus=Cylindropuntia
 
|genus=Cylindropuntia
 
|species=Cylindropuntia fulgida
 
|species=Cylindropuntia fulgida
|abaxial spine length=longest
 
|abaxial spine orientation=spreading;erect;deflexed
 
|abaxial spine shape=flattened
 
|abaxial spine some measurement=0cm;3.5cm
 
|adaxial spine length=longest
 
|adaxial spine orientation=spreading;erect
 
|adaxial spine shape=terete;subterete
 
|adaxial spine some measurement=0cm;2.5cm
 
|anther coloration=white;cream
 
|areole arrangement=adjacent
 
|areole atypical length=7mm;10mm
 
|areole count=18;35
 
|areole length=5mm;7mm
 
|areole shape=obdeltate
 
|areole width=2.5mm;4mm
 
|basal fruit length=32mm;55mm
 
|basal fruit width=23mm;45mm
 
|chain architecture=branching
 
|chain length or size=long
 
|chain orientation=pendent
 
|crown architecture=branched
 
|crown count=many
 
|crown orientation=spreading
 
|depression count=1;2
 
|depression size=large
 
|filament coloration=pale-pink;magenta
 
|fruit architecture=spineless;stipitate
 
|fruit coloration=gray-green
 
|fruit life cycle=proliferating
 
|fruit relief=tuberculate
 
|fruit shape=obconic
 
|fruit texture=fleshy
 
|girdle architecture or pubescence or relief=smooth
 
|glochid arrangement=scattered
 
|glochid coloration=yellow
 
|glochid some measurement=1mm;3mm
 
|hilum shape=pointed
 
|inner tepal architecture=ligulate
 
|inner tepal architecture or shape=emarginate;apiculate
 
|inner tepal coloration=pink;magenta
 
|inner tepal orientation=reflexed
 
|inner tepal shape=obovate
 
|inner tepal some measurement=12mm;16mm
 
|interlaced coloration=brown;pale pinkish;yellowish;brown;pale pinkish;yellowish
 
|interlaced count=absent;absent
 
|interlaced life cycle=aging;aging
 
|seed architecture or course=irregular
 
|seed arrangement or shape=angular
 
|seed coloration=pale-yellow;brownish
 
|seed length=1.9
 
|seed shape=warped
 
|seed width=1.5mm;3.5mm
 
|sheath coloration=yellowish;golden
 
|spine atypical count=12;18
 
|spine count=0;12
 
|stem segment architecture or shape=spiny
 
|stem segment arrangement=subwhorled;whorled
 
|stem segment atypical length=16cm;23cm
 
|stem segment coloration=blackish;gray-green
 
|stem segment condition=drying
 
|stem segment fragility=dislodged
 
|stem segment length=6cm;16cm
 
|stem segment position or structure subtype=terminal
 
|stem segment width=2cm;3.5cm
 
|stigma lobe coloration=whitish;pale-yellow
 
|style coloration=pinkish
 
|terminal fruit length=2cm;3.3cm
 
|terminal fruit width=1.3cm;2.3cm
 
|tree some measurement=1m;3m
 
|trunk architecture=branching
 
|tubercle atypical some measurement=1.3cm;1.9cm
 
|tubercle orientation=salient
 
|tubercle prominence=obscure
 
|tubercle shape=oval
 
|tubercle some measurement=0.8cm;1.3cm
 
|umbilicus depth=deep
 
|umbilicus some measurement=0mm;8mm
 
|wool coloration=gray;black
 
 
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Revision as of 13:59, 27 July 2019

Trees 1–3 m; trunk divaricately branching; crown many branched, spreading. Stem segments whorled or subwhorled, gray-green, often drying blackish, ± spiny throughout, terminal ones easily dislodged, 6–16(–23) × 2–3.5 cm; tubercles salient, broadly oval, 0.8–1.3(–1.9) cm; areoles obdeltate, 5–7(–10) × 2.5–4 mm; wool gold to tan, aging gray to black. Spines 0–12(–18) per areole, at most areoles to nearly absent, yellowish, sometimes also pale pinkish, aging brown, interlaced or not with spines of adjacent areoles; abaxial spines erect to deflexed, spreading, flattened basally, the longest to 3.5 cm; adaxial spines erect or spreading, terete to subterete, longest to 2.5 cm; sheaths uniformly whitish, yellowish to golden, baggy. Glochids in adaxial tuft, sometimes also scattered along areole margins, yellow, 1–3 mm. Flowers: inner tepals usually reflexed, pink to magenta, obovate to ligulate, 12–16 mm, apiculate emarginate; filaments pale pink to magenta; anthers white to cream; style pinkish; stigma lobes whitish to pale yellow. Fruits proliferating, forming long, branching, pendent chains, at maturity gray-green, often stipitate, obconic, fleshy, shallowly tuberculate, usually spineless; basal fruits 32–55 × 23–45 mm; terminal fruits 2–3.3 × 1.3–2.3 cm; tubercles becoming obscure; umbilicus to 8 mm deep; areoles 18–35. Seeds pale yellow to brownish, angular to very irregular in outline, warped, 1.9 × 1.5–3.5 mm, sides with 1–2 large depressions, hilum pointed; girdle smooth.

Distribution

V4 200-distribution-map.gif

sw United States, nw Mexico.

Discussion

Varieties 2 (2 in the flora).

Intermediates are known between the varieties, which are largely sympatric in northern portion of range of the species.

Cylindropuntia fulgida forms hybrids with C. spinosior (see 6. C. ×kelvinensis) and C. leptocaulis. Hybrids, which are rare in south-central Arizona, have stems of intermediate diameter, (0–)1–5 spines per areole, one spine much longer than others, and spineless, yellowing, and often reddish fruits in chains of four to six, or more.

Selected References

None.

Key

1 Spines of stems interlaced with spines of adjacent areoles, the longest usually 2.5-3 cm; sheaths baggy; stem segments appearing spiny from afar, obscuring strongly mammillate tubercles beneath. Cylindropuntia fulgida var. fulgida
1 Spines of stems not or little interlaced with spines of adjacent areoles, the longest usually 1-2 cm; sheaths tightly fitting; stem segments appearing spineless or nearly so from afar, exposing strongly mammillate tubercles beneath Cylindropuntia fulgida var. mamillata