Difference between revisions of "Cimicifuga"

Wernischeck

Gen. Pl., 298. 1763.

Common names: Bugbane snakeroot cohosh
Etymology: Latin cimex, bug, and fugare, to drive away
Treatment appears in FNA Volume 3.
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--><span class="statement" id="st-d0_s0" data-properties="herb duration;rhizome texture;rhizome shape;rhizome duration"><b>Herbs,</b> perennial, from hard, knotted, long-lived rhizomes.</span> <span class="statement" id="st-d0_s1" data-properties="leaf position;leaf position;leaf architecture;leaf architecture;stem architecture or fixation"><b>Leaves </b>basal and cauline, compound, petiolate with basal wings clasping stem;</span> <span class="statement" id="st-d0_s2" data-properties="cauline leaf arrangement">cauline leaves alternate.</span> <span class="statement" id="st-d0_s3" data-properties="leaf-blade architecture"><b>Leaf-</b>blade 1-3-ternately compound;</span> <span class="statement" id="st-d0_s4" data-properties="leaflet shape;leaflet shape;leaflet shape;leaflet shape;leaflet shape;lobe margin shape;lobe margin depth;lobe margin shape">leaflets ovatelanceolate to broadly obovate or orbiculate, 2-5-lobed, lobe margins toothed or shallow to deeply incised.</span> <span class="statement" id="st-d0_s5" data-properties="inflorescence position or structure subtype;panicle architecture;panicle some measurement;branch arrangement"><b>Inflorescences </b>terminal, many-flowered panicles of racemelike branches [spikes in Asian spp.], 7-60 cm;</span> <span class="statement" id="st-d0_s6" data-properties="beak quantity;beak quantity;beak arrangement">bracts 1 or 3, alternate, subtending pedicel (pedicels bracteolate in C. americana), not forming involucre.</span> <span class="statement" id="st-d0_s7" data-properties="flower reproduction;flower architecture or shape"><b>Flowers </b>bisexual [unisexual], radially symmetric;</span> <span class="statement" id="st-d0_s8" data-properties="sepal duration;sepal coloration;sepal coloration;sepal coloration;sepal coloration;sepal coloration;sepal coloration;sepal shape;sepal shape;sepal shape;sepal shape;sepal shape;sepal shape;sepal shape;sepal shape;sepal shape;sepal shape;sepal some measurement;fruit atypical quantity;fruit atypical quantity;fruit quantity">sepals not persistent in fruit, (2-) 4-5 (-6), greenish white or cream to greenish yellow, sometimes pinkish or tinged with red, plane or ± concave, ovate to obovate, 3-6 mm;</span> <span class="statement" id="st-d0_s9" data-properties="petal quantity;petal fusion;petal coloration;petal coloration;petal shape;apex architecture or shape;apex shape;apex some measurement">petals 0-8, distinct, white or yellowish, plane, apex 2-cleft [entire], sometimes clawed, 3-6 mm;</span> <span class="statement" id="st-d0_s10" data-properties="nectariferous area presence">nectariferous area sometimes present;</span> <span class="statement" id="st-d0_s11" data-properties="stamen quantity">stamens 20-110;</span> <span class="statement" id="st-d0_s12" data-properties="filament shape">filaments filiform [flattened];</span> <span class="statement" id="st-d0_s13" data-properties="staminode presence">staminodes absent between stamens and pistils;</span> <span class="statement" id="st-d0_s14" data-properties="pistil quantity;pistil architecture">pistils 1-8, simple;</span> <span class="statement" id="st-d0_s15" data-properties="ovule quantity">ovules 4-15 per pistil;</span> <span class="statement" id="st-d0_s16" data-properties="style presence">style present.</span> <span class="statement" id="st-d0_s17" data-properties="fruits follicle architecture;fruits follicle architecture;fruits follicle architecture;fruits follicle shape;fruits follicle shape;fruits follicle shape;fruits follicle shape;fruits follicle shape;fruits follicle shape;fruits follicle shape;fruits follicle shape;side architecture"><b>Fruits </b>follicles, usually aggregate, sessile or stipitate, ovoid to obovoid, weakly to strongly compressed, sides not prominently veined;</span> <span class="statement" id="st-d0_s18" data-properties="beak position or structure subtype;beak course;beak shape;beak some measurement">beak terminal, straight or hooked at tip, 0.5-2.5 mm.</span> <span class="statement" id="st-d0_s19" data-properties=""><b>Seeds </b>pale-brown to reddish or purplish brown, angled or laterally compressed, hemispheric, lenticular, or cylindric, smooth, slightly ridged, verrucose, or densely scaly.</span> <span class="statement" id="st-d0_s20" data-properties="seed coloration;seed coloration;seed coloration;seed coloration;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed architecture or pubescence or relief;seed shape;seed relief;seed architecture or pubescence;x chromosome quantity">x = 8.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Herbs,</b> perennial, from hard, knotted, long-lived rhizomes. <b>Leaves</b> basal and cauline, compound, petiolate with basal wings clasping stem; cauline leaves alternate. <b>Leaf</b> blade 1-3-ternately compound; leaflets ovate-lanceolate to broadly obovate or orbiculate, 2-5-lobed, lobe margins toothed or shallow to deeply incised. <b>Inflorescences</b> terminal, many-flowered panicles of racemelike branches [spikes in Asian spp.], 7-60 cm; bracts 1 or 3, alternate, subtending pedicel (pedicels bracteolate in C. americana), not forming involucre. <b>Flowers</b> bisexual [unisexual], radially symmetric; sepals not persistent in fruit, (2-)4-5(-6), greenish white or cream to greenish yellow, sometimes pinkish or tinged with red, plane or ± concave, ovate to obovate, 3-6 mm; petals 0-8, distinct, white or yellowish, plane, apex 2-cleft [entire], sometimes clawed, 3-6 mm; nectariferous area sometimes present; stamens 20-110; filaments filiform [flattened]; staminodes absent between stamens and pistils; pistils 1-8, simple; ovules 4-15 per pistil; style present. <b>Fruits</b> follicles, usually aggregate, sessile or stipitate, ovoid to obovoid, weakly to strongly compressed, sides not prominently veined; beak terminal, straight or hooked at tip, 0.5-2.5 mm. <b>Seeds</b> pale brown to reddish or purplish brown, angled or laterally compressed, hemispheric, lenticular, or cylindric, smooth, slightly ridged, verrucose, or densely scaly. <b>x</b> = 8.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
|distribution=North America and Eurasia
+
|distribution=North America and Eurasia.
 
|discussion=<p>Species 12 (6 in the flora).</p><!--
 
|discussion=<p>Species 12 (6 in the flora).</p><!--
 
--><p>Cimicifuga may be divided into two natural groups: those with seeds scaly and those with seeds lacking scales or nearly so. Cimicifuga racemosa and C. elata of North America, with scaleless seeds, are most closely related to C. biternata (Siebold & Zuccarini) Miquel and C. japonica (Thunberg) Sprengel of Asia.</p><!--
 
--><p>Cimicifuga may be divided into two natural groups: those with seeds scaly and those with seeds lacking scales or nearly so. Cimicifuga racemosa and C. elata of North America, with scaleless seeds, are most closely related to C. biternata (Siebold & Zuccarini) Miquel and C. japonica (Thunberg) Sprengel of Asia.</p><!--
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|basionyms=
 
|basionyms=
 
|family=Ranunculaceae
 
|family=Ranunculaceae
|distribution=North America and Eurasia
+
|distribution=North America and Eurasia.
 
|reference=compton1992a;ramsey1965a;ramsey1987a
 
|reference=compton1992a;ramsey1965a;ramsey1987a
 
|publication title=Gen. Pl.,
 
|publication title=Gen. Pl.,
 
|publication year=1763
 
|publication year=1763
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V3/V3_686.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V3/V3_686.xml
 
|genus=Cimicifuga
 
|genus=Cimicifuga
|apex architecture or shape=2-cleft
 
|apex shape=clawed
 
|apex some measurement=3mm;6mm
 
|beak arrangement=alternate
 
|beak course=straight
 
|beak position or structure subtype=terminal
 
|beak quantity=3;1
 
|beak shape=hooked
 
|beak some measurement=0.5mm;2.5mm
 
|branch arrangement=raceme-like
 
|cauline leaf arrangement=alternate
 
|filament shape=filiform
 
|flower architecture or shape=symmetric
 
|flower reproduction=bisexual
 
|fruit atypical quantity=5;6
 
|fruit quantity=4;5
 
|fruits follicle architecture=stipitate;sessile;aggregate
 
|fruits follicle shape=ovoid;obovoid
 
|herb duration=perennial
 
|inflorescence position or structure subtype=terminal
 
|leaf architecture=petiolate;compound
 
|leaf position=cauline;basal
 
|leaf-blade architecture=compound
 
|leaflet shape=2-5-lobed;ovatelanceolate;broadly obovate or orbiculate
 
|lobe margin depth=shallow
 
|lobe margin shape=incised;toothed
 
|nectariferous area presence=absent
 
|ovule quantity=4;15
 
|panicle architecture=many-flowered
 
|panicle some measurement=7cm;60cm
 
|petal coloration=yellowish;white
 
|petal fusion=distinct
 
|petal quantity=0;8
 
|petal shape=plane
 
|pistil architecture=simple
 
|pistil quantity=1;8
 
|rhizome duration=long-lived
 
|rhizome shape=knotted
 
|rhizome texture=hard
 
|seed architecture or pubescence=scaly
 
|seed architecture or pubescence or relief=smooth
 
|seed coloration=pale-brown;reddish or purplish brown
 
|seed relief=verrucose
 
|seed shape=ridged;cylindric;lenticular;hemispheric;compressed;angled;cylindric;lenticular;hemispheric;compressed;angled;cylindric;lenticular;hemispheric;compressed;angled;cylindric;lenticular;hemispheric;compressed;angled
 
|sepal coloration=tinged with red;pinkish;cream;greenish yellow
 
|sepal duration=not persistent
 
|sepal shape=more or less concave ovate;obovate
 
|sepal some measurement=3mm;6mm
 
|side architecture=veined
 
|stamen quantity=20;110
 
|staminode presence=absent
 
|stem architecture or fixation=clasping
 
|style presence=absent
 
|x chromosome quantity=8
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Ranunculaceae]]
 
-->[[Category:Treatment]][[Category:Ranunculaceae]]

Revision as of 13:47, 27 July 2019

Herbs, perennial, from hard, knotted, long-lived rhizomes. Leaves basal and cauline, compound, petiolate with basal wings clasping stem; cauline leaves alternate. Leaf blade 1-3-ternately compound; leaflets ovate-lanceolate to broadly obovate or orbiculate, 2-5-lobed, lobe margins toothed or shallow to deeply incised. Inflorescences terminal, many-flowered panicles of racemelike branches [spikes in Asian spp.], 7-60 cm; bracts 1 or 3, alternate, subtending pedicel (pedicels bracteolate in C. americana), not forming involucre. Flowers bisexual [unisexual], radially symmetric; sepals not persistent in fruit, (2-)4-5(-6), greenish white or cream to greenish yellow, sometimes pinkish or tinged with red, plane or ± concave, ovate to obovate, 3-6 mm; petals 0-8, distinct, white or yellowish, plane, apex 2-cleft [entire], sometimes clawed, 3-6 mm; nectariferous area sometimes present; stamens 20-110; filaments filiform [flattened]; staminodes absent between stamens and pistils; pistils 1-8, simple; ovules 4-15 per pistil; style present. Fruits follicles, usually aggregate, sessile or stipitate, ovoid to obovoid, weakly to strongly compressed, sides not prominently veined; beak terminal, straight or hooked at tip, 0.5-2.5 mm. Seeds pale brown to reddish or purplish brown, angled or laterally compressed, hemispheric, lenticular, or cylindric, smooth, slightly ridged, verrucose, or densely scaly. x = 8.

Distribution

North America and Eurasia.

Discussion

Species 12 (6 in the flora).

Cimicifuga may be divided into two natural groups: those with seeds scaly and those with seeds lacking scales or nearly so. Cimicifuga racemosa and C. elata of North America, with scaleless seeds, are most closely related to C. biternata (Siebold & Zuccarini) Miquel and C. japonica (Thunberg) Sprengel of Asia.

Four or five species of Cimicifuga are cultivated as ornamentals, and at least five named cultivars have been developed.

Key

1 Seeds without scales or scales very short; stigma 0.5 mm wide. > 2
1 Seeds scaly; stigma minute, 0.2–0.3 mm wide. > 3
2 Follicles ovoid; seeds hemispheric, smooth or ±rough-ridged, brown; pedicel subtended by 1 bract; base of terminal leaflet with 3 prominent veins; petals (1–)4(–8), oblong; e North America. Cimicifuga racemosa
2 Follicles oblong; seeds lenticular, usually verrucose, rarely with very short scales, reddish to purplish brown; pedicel subtended by 3 bracts; base of terminal leaflet with 5–7 prominent veins; petals absent; w North America. Cimicifuga elata
3 Pistils sessile, 1–2(–4 in C. arizonica); seeds lenticular, covering of scales giving cylindric appearance. > 4
3 Pistils short-stipitate, (2–)3–8; seeds lenticular, scales not causing cylindric appearance. > 5
4 Petals absent; filaments 3–5 mm; pistils 1(–2), sparsely glandular; style short, straight or slightly recurved; terminal leaflet deeply cordate with 5–9 prominent veins at base; e North America. Cimicifuga rubifolia
4 Petals (0–)2; filaments 5–8 mm; pistils 1–3(–4), glandular; style long, hooked; terminal leaflet somewhat cordate with 3 prominent veins at base; w North America. Cimicifuga arizonica
5 Pistils glabrous; pedicel bracteolate, granular; petiole glabrous; seeds ca. 3.5 mm, covered with broad, lacerate scales; stamens 40–70; filaments 6–10 mm; e North America. Cimicifuga americana
5 Pistils densely pubescent; pedicel not bracteolate; petiole densely pubescent; seeds ca. 2.5–3 mm, loosely covered with narrow, lacerate scales; stamens 20–25; filaments 4–5 mm; w North America. Cimicifuga laciniata