Difference between revisions of "Polypodium glycyrrhiza"

D. C. Eaton

Amer. J. Sci. Arts ser. 2, 22: 138. 1856.

Common names: Licorice fern
Synonyms: Polypodium aleuticum A. E. BobrovPolypodium falcatum KelloggPolypodium occidentale (Hooker) MaxonPolypodium vulgare subsp. occidentale (Hooker) HulténPolypodium vulgare var. falcatum (Kellogg) H. ChristPolypodium vulgare var. occidentale Hooker
Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties="stem coloration;stem coating;stem size;stem size;stem size;stem diameter;stem taste;stem taste"><b>Stems </b>not whitish pruinose, slender to moderately stout, to 6 mm diam., intensely sweet, licorice-flavored;</span> <span class="statement" id="st-d0_s1" data-properties="scale coloration;scale coloration;scale coloration;scale shape;scale shape;scale shape;scale architecture or shape;margin architecture or shape">scales concolored, brown or slightly darker near point of attachment, lanceolate to lanceolate-ovate, symmetric, margins entire.</span> <span class="statement" id="st-d0_s2" data-properties=""><b>Leaves </b>to 75 cm.</span> <span class="statement" id="st-d0_s3" data-properties="petiole size;petiole diameter"><b>Petiole </b>usually slender, 0.5–2 mm diam.</span> <span class="statement" id="st-d0_s4" data-properties="blade shape;blade shape;blade shape;blade shape;blade width;blade width;blade growth form or texture;blade texture"><b>Blade </b>lanceolate-ovate to oblong, pinnatifid, widest near middle or just below, to 16 cm wide, herbaceous, rarely slightly leathery;</span> <span class="statement" id="st-d0_s5" data-properties="rachis pubescence;rachis pubescence;rachis pubescence;rachis pubescence">rachis sparsely scaly to glabrescent abaxially, puberulent adaxially;</span> <span class="statement" id="st-d0_s6" data-properties="scale arrangement or course or shape;scale count;cell width">scales linear, usually less than 3 cells wide.</span> <span class="statement" id="st-d0_s7" data-properties="segment shape;segment shape;segment shape;segment width"><b>Segments </b>linear to oblong, less than 12 mm wide;</span> <span class="statement" id="st-d0_s8" data-properties="margin architecture or shape">margins serrate;</span> <span class="statement" id="st-d0_s9" data-properties="apex shape;apex shape;apex shape">apex acute to attenuate;</span> <span class="statement" id="st-d0_s10" data-properties="">midrib puberulent adaxially.</span> <span class="statement" id="st-d0_s11" data-properties="midrib pubescence;midrib fusion"><b>Venation </b>free.</span> <span class="statement" id="st-d0_s12" data-properties="sorus position;sorus diameter;sorus shape;sorus shape;sorus shape"><b>Sori </b>midway between margin and midrib or slightly closer to midrib, usually less than 3 mm diam., circular to oval when immature.</span> <span class="statement" id="st-d0_s13" data-properties="sporangiaster count"><b>Sporangiasters </b>absent.</span> <span class="statement" id="st-d0_s14" data-properties="spore some measurement;spore relief;surface projection some measurement;2n chromosome count"><b>Spores </b>less than 58 µm, verrucose, with surface projections less than 3 µm. <b>2n</b> = 74.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>not whitish pruinose, slender to moderately stout, to 6 mm diam., intensely sweet, licorice-flavored; scales concolored, brown or slightly darker near point of attachment, lanceolate to lanceolate-ovate, symmetric, margins entire. <b>Leaves</b> to 75 cm. <b>Petiole</b> usually slender, 0.5–2 mm diam. <b>Blade</b> lanceolate-ovate to oblong, pinnatifid, widest near middle or just below, to 16 cm wide, herbaceous, rarely slightly leathery; rachis sparsely scaly to glabrescent abaxially, puberulent adaxially; scales linear, usually less than 3 cells wide. <b>Segments</b> linear to oblong, less than 12 mm wide; margins serrate; apex acute to attenuate; midrib puberulent adaxially. <b>Venation</b> free. <b>Sori</b> midway between margin and midrib or slightly closer to midrib, usually less than 3 mm diam., circular to oval when immature. <b>Sporangiasters</b> absent. <b>Spores</b> less than 58 µm, verrucose, with surface projections less than 3 µm. <b>2n</b> = 74.</span><!--
  
 
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|phenology=Sporulating late fall–spring.
 
|habitat=Cliffs and rocky slopes along coasts, often epiphytic, on a variety of substrates
 
|habitat=Cliffs and rocky slopes along coasts, often epiphytic, on a variety of substrates
 
|elevation=0–700 m.
 
|elevation=0–700 m.
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|basionyms=
 
|basionyms=
 
|family=Polypodiaceae
 
|family=Polypodiaceae
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|phenology=Sporulating late fall–spring.
 
|habitat=Cliffs and rocky slopes along coasts, often epiphytic, on a variety of substrates
 
|habitat=Cliffs and rocky slopes along coasts, often epiphytic, on a variety of substrates
 
|elevation=0–700 m.
 
|elevation=0–700 m.
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|publication year=1856
 
|publication year=1856
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_269.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_269.xml
 
|genus=Polypodium
 
|genus=Polypodium
 
|species=Polypodium glycyrrhiza
 
|species=Polypodium glycyrrhiza
|2n chromosome count=74
 
|apex shape=acute;attenuate
 
|blade growth form or texture=herbaceous
 
|blade shape=pinnatifid;lanceolate-ovate;oblong
 
|blade texture=leathery
 
|blade width=0cm;16cm
 
|cell width=wide
 
|margin architecture or shape=serrate;entire
 
|midrib fusion=free
 
|midrib pubescence=puberulent
 
|petiole diameter=0.5mm;2mm
 
|petiole size=slender
 
|rachis pubescence=puberulent;sparsely scaly;glabrescent abaxially
 
|scale architecture or shape=symmetric
 
|scale arrangement or course or shape=linear
 
|scale coloration=darker;brown;concolored
 
|scale count=0;3
 
|scale shape=lanceolate;lanceolate-ovate
 
|segment shape=linear;oblong
 
|segment width=0mm;12mm
 
|sorus diameter=0mm;3mm
 
|sorus position=midway
 
|sorus shape=circular;oval
 
|sporangiaster count=absent
 
|spore relief=verrucose
 
|spore some measurement=0um;58um
 
|stem coating=pruinose
 
|stem coloration=not whitish
 
|stem diameter=0mm;6mm
 
|stem size=slender;moderately stout
 
|stem taste=licorice-flavored;sweet
 
|surface projection some measurement=0um;3um
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Polypodium]]
 
-->[[Category:Treatment]][[Category:Polypodium]]

Revision as of 13:15, 27 July 2019

Stems not whitish pruinose, slender to moderately stout, to 6 mm diam., intensely sweet, licorice-flavored; scales concolored, brown or slightly darker near point of attachment, lanceolate to lanceolate-ovate, symmetric, margins entire. Leaves to 75 cm. Petiole usually slender, 0.5–2 mm diam. Blade lanceolate-ovate to oblong, pinnatifid, widest near middle or just below, to 16 cm wide, herbaceous, rarely slightly leathery; rachis sparsely scaly to glabrescent abaxially, puberulent adaxially; scales linear, usually less than 3 cells wide. Segments linear to oblong, less than 12 mm wide; margins serrate; apex acute to attenuate; midrib puberulent adaxially. Venation free. Sori midway between margin and midrib or slightly closer to midrib, usually less than 3 mm diam., circular to oval when immature. Sporangiasters absent. Spores less than 58 µm, verrucose, with surface projections less than 3 µm. 2n = 74.


Phenology: Sporulating late fall–spring.
Habitat: Cliffs and rocky slopes along coasts, often epiphytic, on a variety of substrates
Elevation: 0–700 m.

Distribution

V2 269-distribution-map.gif

B.C., Yukon, Alaska, Calif., Idaho, Oreg., Wash., Asia in Kamchatka in the former Soviet republics.

Discussion

Polypodium glycyrrhiza hybridizes with P. calirhiza and with P. hesperium to produce sterile triploids with misshapen spores. Polypodium glycyrrhiza was involved in the origin of both of these allotetraploid species, and some individuals can be difficult to identify. Free versus anastomosing venation distinguishes this species from P. calirhiza; the presence of adaxial hairs on the rachis separates it from P. hesperium. An additional character for distinguishing these taxa is spore length, which is less than 58 µm in diploid P. glycyrrhiza and more than 58 µm in the two tetraploid species. Reports of P. glycyrrhiza occurring in Arizona (T. Reeves 1981; D. B. Lellinger 1985) are based on misidentified specimens.

Selected References

None.

Lower Taxa

None.
... more about "Polypodium glycyrrhiza"
Christopher H. Haufler +, Michael D. Windham +, Frank A. Lang +  and S. A. Whitmore +
D. C. Eaton +
Licorice fern +
B.C. +, Yukon +, Alaska +, Calif. +, Idaho +, Oreg. +, Wash. +  and Asia in Kamchatka in the former Soviet republics. +
0–700 m. +
Cliffs and rocky slopes along coasts, often epiphytic, on a variety of substrates +
Sporulating late fall–spring. +
Amer. J. Sci. Arts +
Polypodium aleuticum +, Polypodium falcatum +, Polypodium occidentale +, Polypodium vulgare subsp. occidentale +, Polypodium vulgare var. falcatum +  and Polypodium vulgare var. occidentale +
Polypodium glycyrrhiza +
Polypodium +
species +