Difference between revisions of "Eriogonum umbellatum"

Revision as of 17:40, 18 September 2019

Herbs, subshrubs, or shrubs, cespitose, matted or spreading, sometimes erect, often polygamo-dioecious, (0.2–)1–12(–20) × (0.5–)1–12(–20) dm, glabrous or tomentose. Stems: caudex spreading; aerial flowering stems spreading to erect or nearly so, slender, solid, not fistulose, arising at nodes of caudex branches and at distal nodes of short, nonflowering aerial branches, (0.1–)0.5–3(–4) dm, without a whorl of bracts at midlength. Leaves in loose to compact basal rosettes; petiole 0.1–3(–4) cm, mostly tomentose to floccose or glabrous; blade oblong-ovate or oblanceolate to elliptic to oval, 0.3–3(–4) × 0.1–2.5 cm, densely lanate to tomentose or floccose abaxially, tomentose to floccose or glabrous adaxially, occasionally glabrous on both surfaces, margins entire, plane or rarely wavy. Inflorescences umbellate or compound-umbellate, rarely subcapitate or capitate, 3–25 × 2–18 cm; branches tomentose to floccose or glabrous, rarely with whorl of bracts ca. midlength; bracts 3–several, semileaflike at proximal node, 0.3–2.5 × 0.2–1.8 cm, usually scalelike distally, 1–5 × 0.5–3 mm. Involucres 1 per node, turbinate to campanulate, 1–6 × (1–)1.5–10 mm, tomentose to thinly floccose or glabrous; teeth 6–12, lobelike, reflexed, 1–4(–6) mm. Flowers 2–10(–12) mm, including (0.7–)1.3–2 mm stipelike base; perianth various shades of white, yellow, or red, glabrous; tepals monomorphic, usually spatulate to obovate; stamens exserted, 2–8 mm; filaments pilose proximally. Achenes light brown to brown, 2–7 mm, glabrous except for sparsely pubescent beak.

w North America.

Discussion

Varieties 41 (41 in the flora).

Eriogonum umbellatum is a widespread and exceedingly variable species rivaling Astragalus lentiginosus Douglas ex Hooker in complexity. Only the variety majus is sometimes recognized at the species rank (as E. subalpinum).

In the following key and descriptions, reference is made to “glabrous” leaf surfaces. This is a function of both age and power of observation. High-power magnification may show some exceedingly fine hairs that are not readily observable to the naked eye. Furthermore, new leaves that ultimately will be “glabrous” will have some fine-tomentose pubescence that becomes less obvious (or even wholly inconspicuous) as the blade expands and matures. Here, the term “glabrous” is used to refer to leaves that are not obviously hairy and are typically bright green on both surfaces when the plant is at full anthesis.

There are several reported uses of sulphur flower in the enthobotany literature, mostly without an indication of the variety. J. B. Romero (1954) indicated that in California, where most of the variants are found, an infusion of the flowers is used for ptomaine poisoning, and M. L. Zigmond (1981) stated that the Kawaiisu used mashed flowers as a salve for gonorrheal sores. E. V. A. Murphey (1959), who worked with P. Train et al. (1941), reported that in Nevada members of the species (most likely var. nevadense) were used in the treatment of colds and stomachaches; J. H. Steward (1933) reported the same uses among the Owens Valley Piute in California. Train and his group noted that poultices of leaves and sometimes roots were used for lameness or rheumatism. In Oregon, leaves of var. ellipticum were used in a poultice to soothe pain, especially that resulting from burns (F. V. Coville 1897; L. Spier 1930). Most of the reports associated with the Navajo or Diné people probably relate to the use of var. subaridum. L. C. Wyman and S. K. Harris (1951) found the species used as a disinfectant or an emetic. The Cheyenne people employed a mixture of powdered stems and flowers to halt lengthy menses (G. B. Grinnell 1923; J. A. Hart 1981). According to A. Johnston (1987), the Blackfoot made a tea from boiled leaves.

The cythera metalmark butterfly (Apodemia mormo cythera) is found in association with a few varieties of sulphur flower (G. F. Pratt and G. R. Ballmer 1991). More commonly seen with the species are the Rocky Mountain dotted-blue (Euphilotes ancilla) and lupine blue (Plebeius lupini).

Selected References

None.

Lower Taxa

Key

1	Inflorescences compound-umbellate, or with at least some branches seemingly with whorl of bracts about midlength	> 2
1	Inflorescences umbellate, not compound-umbellate or with any branches seemingly with whorl of bracts about midlength	> 22
2	Inflorescence branches with a whorl of bracts at midlength (see also E. umbellatum var. polyanthum)	> 3
2	Inflorescence branches without a whorl of bracts at midlength, bracts restricted to base of inflorescence or involucres	> 4
3	Perianths bright yellow; flowers 7-10 mm; Sierra Nevada, c California	Eriogonum umbellatum var. torreyanum
3	Perianths cream or whitish; flowers 4-7 mm; Warner Mountains, se Oregon and ne California	Eriogonum umbellatum var. glaberrimum
4	Flowers 7-10(-12) mm; shrubs 5-15(-20) × 5-20 dm; inflorescences branched 2-4 times; Humboldt and Trinity counties, California	Eriogonum umbellatum var. speciosum
4	Flowers 3-8 mm; matted herbs or subshrubs, or, if shrubby, inflorescences branched 4 or more times, or not of Humboldt and Trinity counties, California	> 5
5	Perianths cream, whitish, or pale yellow to greenish yellow, becoming reddish brown to rose or pink	> 6
5	Perianths bright yellow, not becoming reddish brown to rose or pink	> 7
6	Perianths yellow, becoming reddish brown to rose or pink, with large reddish spot on each midrib; plants spreading to somewhat prostrate mats; se California, s Nevada	Eriogonum umbellatum var. versicolor
6	Perianths cream, whitish, or pale yellow to greenish yellow, without large reddish spot on midrib; plants subshrubs or shrubs; nc Arizona, se California, s Nevada, s Utah	Eriogonum umbellatum var. juniporinum
7	Leaf blades thinly floccose, glabrous, or densely lanate to tomentose on both surfaces at anthesis (see also E. umbellatum var. munzii, s California)	> 8
7	Leaf blades thinly to densely tomentose or lanate abaxially, less so to floccose, nearly glabrous, or glabrous adaxially (rarely both surfaces tomentose in E. umbellatum var. munzii, s California)	> 13
8	Leaf blades densely lanate on both surfaces; wc California	Eriogonum umbellatum var. bahiiforme
8	Leaf blades lanate to tomentose abaxially, tomentose to densely floccose adaxially, sometimes thinly floccose or glabrous on both surfaces; nw or ec California, Intermountain West, or Pacific Northwest.	> 9
9	Leaf blades lanate to tomentose abaxially, thinly tomentose to floccose and greenish adaxially; nw California	> 10
9	Leaf blades usually thinly floccose or glabrous on both surfaces; not of nw California	> 11
10	Leaf blades 0.3-0.7 cm wide; involucral lobes 1-3 mm; plants of serpentine soils; Glenn, Lake, Mendocino, Sonoma, Tehama, and Trinity counties, California	Eriogonum umbellatum var. smallianum
10	Leaf blades (0.5-)0.8-1.8(-2) cm wide; involucral lobes (3-)4-6 mm; plants of nonserpentine soils; Siskiyou County,California	Eriogonum umbellatum var. lautum
11	Leaf blades thinly floccose on both surfaces, or glabrous and green adaxially (rarely glabrous on both surfaces in s Utah, or tomentose abaxially in se Utah); e and s California to sw Colorado and ne Arizona	Eriogonum umbellatum var. subaridum
11	Leaf blades glabrous on both surfaces; e Idaho, ne Oregon, se Washington, and ec California	> 12
12	Herbs; leaf blades broadly elliptic, 1-1.5 cm wide; e Idaho, ne Oregon, and se Washington	Eriogonum umbellatum var. devestivum
12	Subshrubs or shrubs; leaf blades oblanceolate to narrowly elliptic, 0.3-1 cm wide; s Mono, Inyo, and ne Tulare counties, California	Eriogonum umbellatum var. chlorothamnus
13	Plants densely branched, shrubs; w foothills, Sierra Nevada, n California	> 14
13	Plants usually matted herbs or more openly and sparsely branched subshrubs or shrubs; mountains, Sierra Nevada, c and n California, wc Nevada, or Siskiyou and Trinity mountains, nw California (but not on the western foothills), sw Oregon, n Arizona, Colorado, Idaho, Montana, or Washington	> 15
14	Leaf blades white-tomentose abaxially; inflorescences branched 1-2(-3) times, branches thinly floccose or glabrous, central branch sometimes seemingly with a whorl of bracts ca. midlength; Butte, Plumas, and Sierra counties	Eriogonum umbellatum var. polyanthum
14	Leaf blades rusty-lanate to tomentose abaxially; inflorescences branched 3-4 time, branches tomentose to floccose, central branch without a whorl of bracts ca. midlength; Butte County	Eriogonum umbellatum var. ahartii
15	Flowering stems mostly sparsely floccose or glabrous.	> 16
15	Flowering stems usually tomentose or floccose, rarely nearly glabrous at maturity	> 18
16	Leaf blades usually narrowly elliptic, 0.3-1(-1.3) cm wide; plants subshrubs or spreading mats; California, Nevada, Oregon	> 17
16	Leaf blades oblong-ovate to elliptic or oval, 1-1.5(-2) cm wide; plants compact mats; n Arizona	Eriogonum umbellatum var. cognatum
17	Leaf margins plane; flowering stems without a single leaflike bract ca. midlength; Sierra Nevada, e California, wc Nevada	Eriogonum umbellatum var. furcosum
17	Leaf margins often finely wavy; flowering stems often with a single leaflike bract ca. midlength; Siskiyou and Trinity mountains, nw California, sw Oregon	Eriogonum umbellatum var. argus
18	Leaf blades usually broadly elliptic to oval	> 19
18	Leaf blades elliptic	> 20
19	Leaf blades gray abaxially; c Colorado	Eriogonum umbellatum var. ramulosum
19	Leaf blades white abaxially; n California	Eriogonum umbellatum var. lautum
20	Leaf blades 1-1.5 cm wide, usually glabrous adaxially; Idaho, w Montana, Oregon, and e Washington	Eriogonum umbellatum var. ellipticum
20	Leaf blades 0.3-1 cm wide, usually floccose adaxially; California	> 21
21	Leaf blades 0.3-0.7 cm wide; nw California	Eriogonum umbellatum var. smallianum
21	Leaf blades 0.5-1 cm wide; s California	Eriogonum umbellatum var. munzii
22	Perianths usually whitish or cream to red, occasionally yellow	> 23
22	Perianths bright yellow (pale yellow in some individuals of var. vernum in Nevada)	> 27
23	Leaf blades densely lanate on both surfaces; perianths lemon yellow to yellowish red; San Gabriel and San Bernardino mountains, e Los Angeles and w San Bernardino counties, California	Eriogonum umbellatum var. minus
23	Leaf blades tomentose to floccose or glabrous adaxially, sometimes glabrous on both surfaces at full anthesis; perianths not lemon yellow or yellowish red; not of e Los Angeles or w San Bernardino counties, California	> 24
24	Leaf blades glabrous on both surfaces; s Idaho and sw Montana to w Wyoming, ne Nevada, and n Utah	Eriogonum umbellatum var. desereticum
24	Leaf blades densely floccose to tomentose or lanate at least abaxially at full anthesis; widespread.	> 25
25	Leaf blades densely lanate abaxially, glabrous and olive green to bright green adaxially; plants compact mats; perianths cream; Rocky Mountains to Cascade Range	Eriogonum umbellatum var. majus
25	Leaf blades densely floccose to tomentose abaxially, less so or glabrous and green adaxially; plants spreading mats, rarely compact; perianths pale yellow to cream or whitish, rarely greenish white or yellow, becoming reddish brown to rose or pink, with large reddish spots on each midrib; Rocky Mountains to Sierra Nevada.	Eriogonum umbellatumvar. dichrocephalum
26	Perianths pale yellow to cream or whitish, rarely greenish white; leaf blades usually greenish adaxially; se Oregon and e California to w Montana, s Idaho, w Wyoming, c Nevada, and n Utah	Eriogonum umbellatum var. dichrocephalum
26	Perianths yellow, becoming reddish brown to rose or pink, with large reddish spot on each midrib; leaf blades usually reddish adaxially; se California, s Nevada	Eriogonum umbellatum var. versicolor
27	Umbels with branches usually longer than 2.5 cm.	> 28
27	Umbels with branches usually shorter than 2.5 cm (immature specimens should be keyed through the opposing couplet as well)	> 36
28	Leaf blades glabrous on both surfaces at full anthesis, occasionally with some individual blades thinly floccose abaxially	Eriogonum umbellatum var. aureum
28	Leaf blades at least thinly tomentose or lanate abaxially	> 29
29	Leaf blades thinly tomentose to thinly floccose on both surfaces, or glabrous adaxially	> 30
29	Leaf blades densely white-lanate or tomentose abaxially, less so to floccose or glabrous adaxially	> 33
30	Plants matted herbs	> 31
30	Plants subshrubs or shrubs	> 32
31	Leaf blades broadly elliptic to ovate, (0.8-)2-3(-3.5) × (0.7-)1-2(-2.5) cm; s Idaho, ne Nevada, wc Wyoming	Eriogonum umbellatum var. stragulum
31	Leaf blades narrowly elliptic, 0.7-2.5 × 0.3-1 cm; nw Arizona	Eriogonum umbellatum var. mohavense
32	Flowers 4-7 mm; Cali-fornia, n Nevada, se Oregon	Eriogonum umbellatum var. nevadense
32	Flowers (5-)6-10 mm; n Nye County, Nevada	Eriogonum umbellatum var. vernum
33	Subshrubs 3-5 dm; non-serpentine soils; n California and sc Oregon	Eriogonum umbellatum var. dumosum
33	Herbs (0.7-)1-4.5(-5) dm; Rocky Mountains or, if in California and Oregon, often on serpentine soils	> 34
34	Plants usually compact mats; nonserpentine soils; Rocky Mountains, sw Montana and e Idaho to Colorado, s and w Wyo-ming, and n Utah	Eriogonum umbellatum var. umbellatum
34	Plants spreading to prostrate mats; often on serpentine soils; n California and sw Oregon.	> 35
35	Flowering stems 1-2.5(-4) dm; leaf blades 0.5-2(-3.5) cm; flowers 6-8(-9) mm; 400-1700(-2100) m	Eriogonum umbellatum var. goodmanii
35	Flowering stems 0.5-1.5 dm; leaf blades 0.5-1(-1.5) cm; flowers 3-6 mm; 1700-2800 m	Eriogonum umbellatum var. humistratum
36	Plants prostrate, mostly in montane to subalpine or alpine communities.	> 37
36	Plants erect to slightly spreading, not prostrate, usually not in subalpine or alpine communities	> 39
37	Leaf blades glabrous on both surfaces at full anthesis	Eriogonum umbellatum var. porteri
37	Leaf blades tomentose at least abaxially at full anthesis	> 38
38	Leaf blades narrowly elliptic, 0.3-0.6(-1) × 0.2-0.4(-0.6) cm, tomentose abaxially, slightly less so and greenish adaxially; s Sierra Nevada and White Mountains, Mono, Inyo, and Tulare counties, California	Eriogonum umbellatum var. covillei
38	Leaf blades broadly elliptic, 0.5-1.5(-2.5) × 0.5-1.2(-1.5) cm, tomentose or glabrous and olive green adaxially; n Cascade Range, Benton, Clackamas, Hood River, and Wasco counties, Oregon, and Kittitas and Yakima counties, Washington	Eriogonum umbellatum var. haussknechtii
39	Leaf blades glabrous on both surfaces, sometimes with marginal hairs at full anthesis	> 40
39	Leaf blades lanate to tomentose or floccose at least abaxially at full anthesis.	> 41
40	Leaf blades without marginal hairs; se Oregon, Idaho, and Nevada to w Montana, Wyoming, Colorado, and Utah	Eriogonum umbellatum var. aureum
40	Leaf blades with marginal hairs; Cascade Range, Washington	Eriogonum umbellatum var. hypoleium
41	Leaf blades thinly tomentose abaxially, less so to floccose or glabrous and green adaxially; Sierra Nevada and Great Basin ranges	Eriogonum umbellatum var. nevadense
41	Leaf blades densely lanate to tomentose abaxially; widespread, generally not in Great Basin ranges.	> 42
42	Mature leaf blades lanate to tomentose on both surfaces; Yellowstone National Park, Wyoming	Eriogonum umbellatum var. cladophorum
42	Mature leaf blades densely lanate or tomentose abaxially, less so to floccose or glabrous and greenish adaxially; w United States	> 43
43	Larger leaf blades usually 0.3-1.5(-2) cm, usually elliptic to oblong; plants subshrubs or herbs	> 44
43	Larger leaf blades 1-3(-3.5) cm, elliptic to ovate; plants subshrubs, or compact or spreading mats	> 45
44	Leaf blades usually elliptic; non-serpentine soils; ne California, se Oregon, sw Idaho, n Nevada	Eriogonum umbellatum var. modocense
44	Leaf blades elliptic to oblong; serpentine outcrops; nw California 107aa. Eriogonum umbellatum var. nelsoniorum	> 43
45	Plants subshrubs; n Cascade Range, Washington	Eriogonum umbellatum var. sandbergii
45	Plants compact or spreading mats; Rocky Mountains, Sierra Nevada, Transverse, and Argus ranges, California, Colorado, Idaho, Montana, Utah, and Wyoming	> 46
46	Leaf blades elliptic to ovate, densely white-lanate abaxially; Rocky Mountains, sw Montana and e Idaho to s Colorado, w and s Wyoming, and s Utah	Eriogonum umbellatum var. umbellatum
46	Leaf blades elliptic, densely grayish-lanate abaxially; s Sierra Nevada, Transverse Ranges, and Argus Range, California	Eriogonum umbellatum var. canifolium

@@ Line 22: / Line 22: @@
 |distribution=w North America.
 |discussion=<p>Varieties 41 (41 in the flora).</p><!--
---><p>Eriogonum umbellatum is a widespread and exceedingly variable species rivaling Astragalus lentiginosus Douglas ex Hooker in complexity. Only the variety majus is sometimes recognized at the species rank (as E. subalpinum).</p><!--
+--><p><i>Eriogonum umbellatum</i> is a widespread and exceedingly variable species rivaling Astragalus lentiginosus Douglas ex Hooker in complexity. Only the variety majus is sometimes recognized at the species rank (as E. subalpinum).</p><!--
 --><p>In the following key and descriptions, reference is made to “glabrous” leaf surfaces. This is a function of both age and power of observation. High-power magnification may show some exceedingly fine hairs that are not readily observable to the naked eye. Furthermore, new leaves that ultimately will be “glabrous” will have some fine-tomentose pubescence that becomes less obvious (or even wholly inconspicuous) as the blade expands and matures. Here, the term “glabrous” is used to refer to leaves that are not obviously hairy and are typically bright green on both surfaces when the plant is at full anthesis.</p><!--
---><p>There are several reported uses of sulphur flower in the enthobotany literature, mostly without an indication of the variety. J. B. Romero (1954) indicated that in California, where most of the variants are found, an infusion of the flowers is used for ptomaine poisoning, and M. L. Zigmond (1981) stated that the Kawaiisu used mashed flowers as a salve for gonorrheal sores. E. V. A. Murphey (1959), who worked with P. Train et al. (1941), reported that in Nevada members of the species (most likely var. nevadense) were used in the treatment of colds and stomachaches; J. H. Steward (1933) reported the same uses among the Owens Valley Piute in California. Train and his group noted that poultices of leaves and sometimes roots were used for lameness or rheumatism. In Oregon, leaves of var. ellipticum were used in a poultice to soothe pain, especially that resulting from burns (F. V. Coville 1897; L. Spier 1930). Most of the reports associated with the Navajo or Diné people probably relate to the use of var. subaridum. L. C. Wyman and S. K. Harris (1951) found the species used as a disinfectant or an emetic. The Cheyenne people employed a mixture of powdered stems and flowers to halt lengthy menses (G. B. Grinnell 1923; J. A. Hart 1981). According to A. Johnston (1987), the Blackfoot made a tea from boiled leaves.</p><!--
+--><p>There are several reported uses of sulphur flower in the enthobotany literature, mostly without an indication of the variety. J. B. Romero (1954) indicated that in California, where most of the variants are found, an infusion of the flowers is used for ptomaine poisoning, and M. L. Zigmond (1981) stated that the Kawaiisu used mashed flowers as a salve for gonorrheal sores. E. V. A. Murphey (1959), who worked with P. Train et al. (1941), reported that in <i>Nevada</i> members of the species (most likely <i></i>var.<i> nevadense</i>) were used in the treatment of colds and stomachaches; J. H. Steward (1933) reported the same uses among the Owens Valley Piute in California. Train and his group noted that poultices of leaves and sometimes roots were used for lameness or rheumatism. In Oregon, leaves of <i></i>var.<i> ellipticum</i> were used in a poultice to soothe pain, especially that resulting from burns (F. V. Coville 1897; L. Spier 1930). Most of the reports associated with the Navajo or Diné people probably relate to the use of <i></i>var.<i> subaridum</i>. L. C. Wyman and S. K. Harris (1951) found the species used as a disinfectant or an emetic. The Cheyenne people employed a mixture of powdered stems and flowers to halt lengthy menses (G. B. Grinnell 1923; J. A. Hart 1981). According to A. Johnston (1987), the Blackfoot made a tea from boiled leaves.</p><!--
 --><p>The cythera metalmark butterfly (Apodemia mormo cythera) is found in association with a few varieties of sulphur flower (G. F. Pratt and G. R. Ballmer 1991). More commonly seen with the species are the Rocky Mountain dotted-blue (Euphilotes ancilla) and lupine blue (Plebeius lupini).</p>
 |tables=
@@ Line 420: / Line 420: @@
 |publication year=1827
 |special status=
-|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V5/V5_673.xml
+|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/8f726806613d60c220dc4493de13607dd3150896/coarse_grained_fna_xml/V5/V5_673.xml
 |subfamily=Polygonaceae subfam. Eriogonoideae
 |genus=Eriogonum