Difference between revisions of "Dryopteris intermedia"

(Muhlenberg ex Willdenow) A. Gray

Manual 630. 1848.

Common names: Evergreen wood fern fancy fern dryoptère spinuleuse
Basionym: Aspidium intermedium Muhlenberg ex Willdenow
Synonyms: Dryopteris austriaca var. intermedia (Muhlenberg ex Willdenow) C. V. MortonDryopteris spinulosa var. intermedia (Muhlenberg ex Willdenow) L. Underwood
Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties="leaf architecture;leaf coloration;leaf length;leaf width"><b>Leaves </b>monomorphic, green through winter, 32–90 × 10–20 cm.</span> <span class="statement" id="st-d0_s1" data-properties="petiole length;petiole architecture or pubescence"><b>Petiole </b>1/3 length of leaf, scaly at least at base;</span> <span class="statement" id="st-d0_s2" data-properties="scale arrangement;scale coloration">scales scattered, tan.</span> <span class="statement" id="st-d0_s3" data-properties="blade coloration;blade shape;blade shape;blade growth form or texture;blade architecture or function or pubescence"><b>Blade </b>green, ovate, 3-pinnate-pinnatifid, herbaceous, glandular.</span> <span class="statement" id="st-d0_s4" data-properties="pinna shape"><b>Pinnae </b>± in plane of blade, lanceolate-oblong;</span> <span class="statement" id="st-d0_s5" data-properties="basal pinna shape;basal pinna size;basal pinnule length or size;pinnule arrangement;pinnule orientation;pinnule length or size;basal pinnule orientation">basal pinnae lanceolate, not reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule longer than basal acroscopic pinnule;</span> <span class="statement" id="st-d0_s6" data-properties="pinnule margin architecture or shape;tooth architecture or shape">pinnule margins serrate, teeth spiny.</span> <span class="statement" id="st-d0_s7" data-properties="sorus position"><b>Sori </b>midway between midvein and margin of segments.</span> <span class="statement" id="st-d0_s8" data-properties="glandular-hair size"><b>Indusia </b>with minute glandular-hairs.</span> <span class="statement" id="st-d0_s9" data-properties="2n chromosome count">2n = 82.</span><!--
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--><span class="statement" id="st-undefined" data-properties=""><b>Leaves </b>monomorphic, green through winter, 32–90 × 10–20 cm. <b>Petiole</b> 1/3 length of leaf, scaly at least at base; scales scattered, tan. <b>Blade</b> green, ovate, 3-pinnate-pinnatifid, herbaceous, glandular. <b>Pinnae</b> ± in plane of blade, lanceolate-oblong; basal pinnae lanceolate, not reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule longer than basal acroscopic pinnule; pinnule margins serrate, teeth spiny. <b>Sori</b> midway between midvein and margin of segments. <b>Indusia</b> with minute glandular hairs. <b>2n</b> = 82.</span><!--
  
 
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|publication year=1848
 
|publication year=1848
 
|special status=
 
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_508.xml
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_508.xml
 
|genus=Dryopteris
 
|genus=Dryopteris
 
|species=Dryopteris intermedia
 
|species=Dryopteris intermedia
|2n chromosome count=82
 
|basal pinna shape=lanceolate
 
|basal pinna size=not reduced
 
|basal pinnule length or size=longer
 
|basal pinnule orientation=acroscopic
 
|blade architecture or function or pubescence=glandular
 
|blade coloration=green
 
|blade growth form or texture=herbaceous
 
|blade shape=3-pinnate-pinnatifid;ovate
 
|glandular-hair size=minute
 
|leaf architecture=monomorphic
 
|leaf coloration=green
 
|leaf length=32cm;90cm
 
|leaf width=10cm;20cm
 
|petiole architecture or pubescence=scaly
 
|petiole length=1/3 length of leaf
 
|pinna shape=lanceolate-oblong
 
|pinnule arrangement=adjacent
 
|pinnule length or size=longer
 
|pinnule margin architecture or shape=serrate
 
|pinnule orientation=basiscopic
 
|scale arrangement=scattered
 
|scale coloration=tan
 
|sorus position=midway
 
|tooth architecture or shape=spiny
 
 
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Revision as of 13:22, 27 July 2019

Leaves monomorphic, green through winter, 32–90 × 10–20 cm. Petiole 1/3 length of leaf, scaly at least at base; scales scattered, tan. Blade green, ovate, 3-pinnate-pinnatifid, herbaceous, glandular. Pinnae ± in plane of blade, lanceolate-oblong; basal pinnae lanceolate, not reduced, basal pinnules longer than adjacent pinnules, basal basiscopic pinnule longer than basal acroscopic pinnule; pinnule margins serrate, teeth spiny. Sori midway between midvein and margin of segments. Indusia with minute glandular hairs. 2n = 82.


Habitat: Moist rocky woods, especially hemlock hardwoods, ravines, and edges of swamps
Elevation: 0–2000 m

Distribution

V2 508-distribution-map.gif

N.B., Nfld., N.S., Ont., P.E.I., Que., Ala., Conn., Del., Ga., Ill., Ind., Iowa, Ky., Maine, Md., Mass., Mich., Minn., Mo., N.H., N.J., N.Y., N.C., Ohio, Pa., R.I., Tenn., Vt., Va., W.Va., Wis.

Discussion

A related taxon, Dryopteris intermedia subsp. maderensis (J. Milde ex Alston) Fraser-Jenkins, occurs on eastern Atlantic islands.

Dryopteris intermedia and the other taxa in the " D. spinulosa complex" have long confounded taxonomists. Dryopteris intermedia is diploid and is one of the parents of the allotetraploids D. carthusiana and D. campyloptera. Dryopteris intermedia hybridizes with eight species. All hybrids are easily detected by the distinctive glandular hairs on the indusia and, usually, on the costae and costules.

Selected References

None.

Lower Taxa

None.
... more about "Dryopteris intermedia"
James D. Montgomery +  and Warren H. Wagner Jr. +
(Muhlenberg ex Willdenow) A. Gray +
Aspidium intermedium +
Evergreen wood fern +, fancy fern +  and dryoptère spinuleuse +
N.B. +, Nfld. +, N.S. +, Ont. +, P.E.I. +, Que. +, Ala. +, Conn. +, Del. +, Ga. +, Ill. +, Ind. +, Iowa +, Ky. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Mo. +, N.H. +, N.J. +, N.Y. +, N.C. +, Ohio +, Pa. +, R.I. +, Tenn. +, Vt. +, Va. +, W.Va. +  and Wis. +
0–2000 m +
Moist rocky woods, especially hemlock hardwoods, ravines, and edges of swamps +
Dryopteris austriaca var. intermedia +  and Dryopteris spinulosa var. intermedia +
Dryopteris intermedia +
Dryopteris +
species +