Difference between revisions of "Cystopteris reevesiana"

Lellinger

Amer. Fern J. 71: 92. 1981.

Common names: Southwestern brittle fern
Synonyms: Cystopteris fragilis subsp. tenuifolia Clute
Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties="stem growth form or orientation;stem shape;internode length or size;petiole base arrangement;petiole base duration;hair count"><b>Stems </b>creeping, not cordlike, internodes usually long, with scattered persistent petiole bases, hairs absent;</span> <span class="statement" id="st-d0_s1" data-properties="scale coloration;scale coloration;scale coloration;scale shape;scale shape;scale shape;wall arrangement;wall width;lumina coloration">scales tan to brown, ovate to lanceolate, radial walls thin, luminae tan.</span> <span class="statement" id="st-d0_s2" data-properties="leaf architecture;leaf arrangement or growth form;leaf some measurement"><b>Leaves </b>monomorphic, clustered at stem apex, to 45 cm, bearing sori throughout year.</span> <span class="statement" id="st-d0_s3" data-properties="petiole color;petiole coloration;petiole height or length or size;blade coloration;blade coloration;blade coloration;blade coloration;blade coloration;blade height or length or size;blade architecture or pubescence;base coloration;base coloration;base coloration;base coloration;base coloration;base height or length or size;base architecture or pubescence;junction architecture or pubescence;blade architecture or pubescence;blade coloration;blade coloration;blade coloration;blade coloration;blade architecture or pubescence;base coloration;base coloration;base coloration;base coloration;base architecture or pubescence"><b>Petiole </b>highly variable in color, from uniformly dark purple to uniformly straw-colored, but mostly dark purple at base, grading to straw-colored at junction with blade, shorter than blade, base sparsely scaly.</span> <span class="statement" id="st-d0_s4" data-properties="blade shape;blade shape;blade shape;blade architecture or shape;blade width;middle position;apex shape"><b>Blade </b>ovate to elliptic, 2–3-pinnate, widest at or just below middle, apex short-attenuate;</span> <span class="statement" id="st-d0_s5" data-properties="hair count;hair architecture;bulblet count;bulblet architecture">rachis and costae lacking gland-tipped hairs or bulblets;</span> <span class="statement" id="st-d0_s6" data-properties="hair count;hair architecture;hair architecture">axils of pinnae with occasional multicellular, gland-tipped hairs.</span> <span class="statement" id="st-d0_s7" data-properties="pinna orientation;pinna course;margin shape;margin shape;margin shape"><b>Pinnae </b>usually perpendicular to rachis, not curving toward blade apex, margins dentate to crenate;</span> <span class="statement" id="st-d0_s8" data-properties="proximal pinna shape;proximal pinna architecture or shape;pinnule orientation;pinnule size">proximal pinnae pinnate-pinnatifid to 2-pinnate, ± equilateral, basiscopic pinnules not enlarged;</span> <span class="statement" id="st-d0_s9" data-properties="pinnule orientation;pinnule architecture;base shape;base shape;base shape;distal pinna shape;distal pinna shape;distal pinna shape">basal basiscopic pinnules mostly short-stalked, base truncate to obtuse, distal pinnae deltate to ovate.</span> <span class="statement" id="st-d0_s10" data-properties="vein orientation"><b>Veins </b>directed into teeth and notches.</span> <span class="statement" id="st-d0_s11" data-properties="indusium shape;indusium shape;indusium shape;hair architecture;hair count"><b>Indusia </b>cupshaped to lanceolate, gland-tipped hairs absent.</span> <span class="statement" id="st-d0_s12" data-properties="spore architecture or shape;2n chromosome count"><b>Spores </b>spiny, usually averaging 33–41 µm. <b>2n</b> = 84.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>creeping, not cordlike, internodes usually long, with scattered persistent petiole bases, hairs absent; scales tan to brown, ovate to lanceolate, radial walls thin, luminae tan. <b>Leaves</b> monomorphic, clustered at stem apex, to 45 cm, bearing sori throughout year. <b>Petiole</b> highly variable in color, from uniformly dark purple to uniformly straw-colored, but mostly dark purple at base, grading to straw-colored at junction with blade, shorter than blade, base sparsely scaly. <b>Blade</b> ovate to elliptic, 2–3-pinnate, widest at or just below middle, apex short-attenuate; rachis and costae lacking gland-tipped hairs or bulblets; axils of pinnae with occasional multicellular, gland-tipped hairs. <b>Pinnae</b> usually perpendicular to rachis, not curving toward blade apex, margins dentate to crenate; proximal pinnae pinnate-pinnatifid to 2-pinnate, ± equilateral, basiscopic pinnules not enlarged; basal basiscopic pinnules mostly short-stalked, base truncate to obtuse, distal pinnae deltate to ovate. <b>Veins</b> directed into teeth and notches. <b>Indusia</b> cup-shaped to lanceolate, gland-tipped hairs absent. <b>Spores</b> spiny, usually averaging 33–41 µm. <b>2n</b> = 84.</span><!--
  
 
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|phenology=Sporulating summer–fall.
 
|habitat=Terrestrial or on rock on variety of substrates
 
|habitat=Terrestrial or on rock on variety of substrates
 
|elevation=1500–4000 m
 
|elevation=1500–4000 m
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|basionyms=
 
|basionyms=
 
|family=Dryopteridaceae
 
|family=Dryopteridaceae
 +
|phenology=Sporulating summer–fall.
 
|habitat=Terrestrial or on rock on variety of substrates
 
|habitat=Terrestrial or on rock on variety of substrates
 
|elevation=1500–4000 m
 
|elevation=1500–4000 m
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|publication year=1981
 
|publication year=1981
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_369.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_369.xml
 
|genus=Cystopteris
 
|genus=Cystopteris
 
|species=Cystopteris reevesiana
 
|species=Cystopteris reevesiana
|2n chromosome count=84
 
|apex shape=short-attenuate
 
|base architecture or pubescence=scaly;scaly
 
|base coloration=dark purple;uniformly dark purple;uniformly straw-colored
 
|base height or length or size=from uniformly dark purple to uniformly straw-colored , but mostly dark purple at base , grading to straw-colored at junction with blade , shorter than blade , base
 
|base shape=truncate;obtuse
 
|blade architecture or pubescence=scaly;scaly;scaly
 
|blade architecture or shape=2-3-pinnate
 
|blade coloration=dark purple;uniformly dark purple;uniformly straw-colored
 
|blade height or length or size=from uniformly dark purple to uniformly straw-colored , but mostly dark purple at base , grading to straw-colored at junction with blade , shorter than blade , base
 
|blade shape=ovate;elliptic
 
|blade width=widest
 
|bulblet architecture=gland-tipped
 
|bulblet count=lacking
 
|distal pinna shape=deltate;ovate
 
|hair architecture=gland-tipped;gland-tipped;multicellular;gland-tipped
 
|hair count=absent;occasional;lacking;absent
 
|indusium shape=cupshaped;lanceolate
 
|internode length or size=long
 
|junction architecture or pubescence=scaly
 
|leaf architecture=monomorphic
 
|leaf arrangement or growth form=clustered
 
|leaf some measurement=0cm;45cm
 
|lumina coloration=tan
 
|margin shape=dentate;crenate
 
|middle position=middle
 
|petiole base arrangement=scattered
 
|petiole base duration=persistent
 
|petiole color=variable
 
|petiole coloration=from uniformly dark purple to uniformly straw-colored , but mostly dark purple at base , grading to straw-colored at junction with blade , shorter than blade , base
 
|petiole height or length or size=from uniformly dark purple to uniformly straw-colored , but mostly dark purple at base , grading to straw-colored at junction with blade , shorter than blade , base
 
|pinna course=not curving
 
|pinna orientation=perpendicular
 
|pinnule architecture=short-stalked
 
|pinnule orientation=basiscopic;basiscopic
 
|pinnule size=not enlarged
 
|proximal pinna architecture or shape=equilateral
 
|proximal pinna shape=pinnate-pinnatifid to 2-pinnate
 
|scale coloration=tan;brown
 
|scale shape=ovate;lanceolate
 
|spore architecture or shape=spiny
 
|stem growth form or orientation=creeping
 
|stem shape=not cordlike
 
|vein orientation=directed
 
|wall arrangement=radial
 
|wall width=thin
 
 
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-->[[Category:Treatment]][[Category:Cystopteris]]
 
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Revision as of 13:18, 27 July 2019

Stems creeping, not cordlike, internodes usually long, with scattered persistent petiole bases, hairs absent; scales tan to brown, ovate to lanceolate, radial walls thin, luminae tan. Leaves monomorphic, clustered at stem apex, to 45 cm, bearing sori throughout year. Petiole highly variable in color, from uniformly dark purple to uniformly straw-colored, but mostly dark purple at base, grading to straw-colored at junction with blade, shorter than blade, base sparsely scaly. Blade ovate to elliptic, 2–3-pinnate, widest at or just below middle, apex short-attenuate; rachis and costae lacking gland-tipped hairs or bulblets; axils of pinnae with occasional multicellular, gland-tipped hairs. Pinnae usually perpendicular to rachis, not curving toward blade apex, margins dentate to crenate; proximal pinnae pinnate-pinnatifid to 2-pinnate, ± equilateral, basiscopic pinnules not enlarged; basal basiscopic pinnules mostly short-stalked, base truncate to obtuse, distal pinnae deltate to ovate. Veins directed into teeth and notches. Indusia cup-shaped to lanceolate, gland-tipped hairs absent. Spores spiny, usually averaging 33–41 µm. 2n = 84.


Phenology: Sporulating summer–fall.
Habitat: Terrestrial or on rock on variety of substrates
Elevation: 1500–4000 m

Distribution

V2 369-distribution-map.gif

Ariz., Colo., N.Mex., Tex., Utah, Mexico.

Discussion

The finely dissected leaves, dark petioles, creeping stems, smaller spores, and terrestrial habit distinguish Cystopteris reevesiana from C. fragilis in the southwest. On rock and at high elevations, however, C. reevesiana can have stems with short internodes and leaves that are reduced in size and dissection (resembling C. fragilis). In southern Colorado, the two species are sympatric in some areas and form triploid hybrids. Cystopteris reevesiana and C. bulbifera are the diploid progenitors of C. utahensis, which occasionally crosses with C. reevesiana to produce sterile triploid hybrids of intermediate morphology.

Selected References

None.

Lower Taxa

None.
... more about "Cystopteris reevesiana"
Christopher H. Haufler +, Robbin C. Moran +  and Michael D. Windham +
Lellinger +
Southwestern brittle fern +
Ariz. +, Colo. +, N.Mex. +, Tex. +, Utah +  and Mexico. +
1500–4000 m +
Terrestrial or on rock on variety of substrates +
Sporulating summer–fall. +
Amer. Fern J. +
Cystopteris fragilis subsp. tenuifolia +
Cystopteris reevesiana +
Cystopteris +
species +