Difference between revisions of "Woodsia neomexicana"

Windham

Contr. Univ. Michigan Herb. 19: 52. 1993.

Common names: New Mexican cliff fern
Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties="stem architecture or arrangement;stem orientation;stem orientation;stem orientation;petiole base count;petiole base duration"><b>Stems </b>compact, erect to ascending, with few-to-many persistent petiole bases of unequal lengths;</span> <span class="statement" id="st-d0_s1" data-properties="scale coloration;scale coloration;scale shape;central stripe coloration;margin coloration">scales mostly uniformly brown but at least some bicolored with dark central stripe and pale-brown margins, narrowly lanceolate.</span> <span class="statement" id="st-d0_s2" data-properties="leaf length;leaf width"><b>Leaves </b>4–30 × 1.5–6 cm.</span> <span class="statement" id="st-d0_s3" data-properties="petiole coloration;petiole coloration;petiole coloration;petiole architecture;petiole fragility"><b>Petiole </b>light-brown or straw-colored when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered.</span> <span class="statement" id="st-d0_s4" data-properties="blade shape;blade shape;blade shape;blade shape;blade pubescence;blade pubescence;blade pubescence;blade coating"><b>Blade </b>linear to lanceolate, usually pinnate-pinnatifid proximally, glabrescent to sparsely glandular, never viscid;</span> <span class="statement" id="st-d0_s5" data-properties="stalk width;tip size">glandular-hairs with thin stalks and slightly expanded tips;</span> <span class="statement" id="st-d0_s6" data-properties="glandular-hair arrangement;scale count;scale shape">rachis with scattered glandular-hairs and rare, hairlike scales.</span> <span class="statement" id="st-d0_s7" data-properties="pinna shape;pinna shape;pinna shape;pinna length or size;pinna shape;apex shape;apex shape"><b>Pinnae </b>ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex;</span> <span class="statement" id="st-d0_s8" data-properties="pair count;pinnule arrangement">largest pinnae with 3–7 pairs of closely spaced pinnules;</span> <span class="statement" id="st-d0_s9" data-properties="abaxial and adaxial surface pubescence;abaxial and adaxial surface architecture or function or pubescence;hair count;hair architecture;scale count;scale architecture">abaxial and adaxial surfaces glabrescent to sparsely glandular, lacking nonglandular hairs or scales.</span> <span class="statement" id="st-d0_s10" data-properties="pinnule architecture or shape;pinnule shape"><b>Pinnules </b>dentate, often shallowly lobed;</span> <span class="statement" id="st-d0_s11" data-properties="margin reflectance;margin width;gland count;cilium count;projection architecture;projection coloration or reflectance">margins nonlustrous, thin, with occasional glands, lacking cilia, with 1–2-celled translucent projections on teeth.</span> <span class="statement" id="st-d0_s12" data-properties="vein tip size;hydathode coloration;hydathode prominence"><b>Vein </b>tips occasionally enlarged to form whitish hydathodes visible adaxially.</span> <span class="statement" id="st-d0_s13" data-properties="indusium length;segment size or width;segment texture;cell architecture or shape;sporangium life cycle"><b>Indusia </b>of narrow, filamentous segments, these uniseriate for most of length, composed of ± isodiametric cells, usually surpassing mature sporangia.</span> <span class="statement" id="st-d0_s14" data-properties="spore some measurement;2n chromosome count"><b>Spores </b>averaging 44–52 µm. <b>2n</b> = 152.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact, erect to ascending, with few to many persistent petiole bases of unequal lengths; scales mostly uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. <b>Leaves</b> 4–30 × 1.5–6 cm. <b>Petiole</b> light brown or straw-colored when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered. <b>Blade</b> linear to lanceolate, usually pinnate-pinnatifid proximally, glabrescent to sparsely glandular, never viscid; glandular hairs with thin stalks and slightly expanded tips; rachis with scattered glandular hairs and rare, hairlike scales. <b>Pinnae</b> ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex; largest pinnae with 3–7 pairs of closely spaced pinnules; abaxial and adaxial surfaces glabrescent to sparsely glandular, lacking nonglandular hairs or scales. <b>Pinnules</b> dentate, often shallowly lobed; margins nonlustrous, thin, with occasional glands, lacking cilia, with 1–2-celled translucent projections on teeth. <b>Vein</b> tips occasionally enlarged to form whitish hydathodes visible adaxially. <b>Indusia</b> of narrow, filamentous segments, these uniseriate for most of length, composed of ± isodiametric cells, usually surpassing mature sporangia. <b>Spores</b> averaging 44–52 µm. <b>2n</b> = 152.</span><!--
  
 
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|phenology=Sporulating summer–fall.
 
|habitat=Cliffs and rocky slopes, usually on sandstone or igneous substrates
 
|habitat=Cliffs and rocky slopes, usually on sandstone or igneous substrates
 
|elevation=300–3500 m
 
|elevation=300–3500 m
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|basionyms=
 
|basionyms=
 
|family=Dryopteridaceae
 
|family=Dryopteridaceae
 +
|phenology=Sporulating summer–fall.
 
|habitat=Cliffs and rocky slopes, usually on sandstone or igneous substrates
 
|habitat=Cliffs and rocky slopes, usually on sandstone or igneous substrates
 
|elevation=300–3500 m
 
|elevation=300–3500 m
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|publication year=1993
 
|publication year=1993
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_245.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_245.xml
 
|genus=Woodsia
 
|genus=Woodsia
 
|species=Woodsia neomexicana
 
|species=Woodsia neomexicana
|2n chromosome count=152
 
|abaxial and adaxial surface architecture or function or pubescence=glandular
 
|abaxial and adaxial surface pubescence=glabrescent to sparsely
 
|apex shape=acute;rounded
 
|blade coating=viscid
 
|blade pubescence=glabrescent;sparsely glandular
 
|blade shape=pinnate-pinnatifid;linear;lanceolate
 
|cell architecture or shape=isodiametric
 
|central stripe coloration=dark
 
|cilium count=lacking
 
|gland count=occasional
 
|glandular-hair arrangement=scattered
 
|hair architecture=nonglandular
 
|hair count=lacking
 
|hydathode coloration=whitish
 
|hydathode prominence=visible
 
|indusium length=uniseriate
 
|leaf length=4cm;30cm
 
|leaf width=1.5cm;6cm
 
|margin coloration=pale-brown
 
|margin reflectance=nonlustrous
 
|margin width=thin
 
|pair count=3;7
 
|petiole architecture=not articulate
 
|petiole base count=few-to-many
 
|petiole base duration=persistent
 
|petiole coloration=darker;straw-colored;light-brown
 
|petiole fragility=brittle
 
|pinna length or size=longer than wide
 
|pinna shape=tapered;ovate-deltate;elliptic
 
|pinnule architecture or shape=dentate
 
|pinnule arrangement=spaced
 
|pinnule shape=lobed
 
|projection architecture=1-2-celled
 
|projection coloration or reflectance=translucent
 
|scale architecture=nonglandular
 
|scale coloration=bicolored;brown
 
|scale count=lacking;rare
 
|scale shape=hairlike;lanceolate
 
|segment size or width=narrow
 
|segment texture=filamentous
 
|sporangium life cycle=mature
 
|spore some measurement=44um;52um
 
|stalk width=thin
 
|stem architecture or arrangement=compact
 
|stem orientation=erect;ascending
 
|tip size=expanded
 
|vein tip size=enlarged
 
 
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-->[[Category:Treatment]][[Category:Woodsia]]
 
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Revision as of 13:15, 27 July 2019

Stems compact, erect to ascending, with few to many persistent petiole bases of unequal lengths; scales mostly uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate. Leaves 4–30 × 1.5–6 cm. Petiole light brown or straw-colored when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered. Blade linear to lanceolate, usually pinnate-pinnatifid proximally, glabrescent to sparsely glandular, never viscid; glandular hairs with thin stalks and slightly expanded tips; rachis with scattered glandular hairs and rare, hairlike scales. Pinnae ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex; largest pinnae with 3–7 pairs of closely spaced pinnules; abaxial and adaxial surfaces glabrescent to sparsely glandular, lacking nonglandular hairs or scales. Pinnules dentate, often shallowly lobed; margins nonlustrous, thin, with occasional glands, lacking cilia, with 1–2-celled translucent projections on teeth. Vein tips occasionally enlarged to form whitish hydathodes visible adaxially. Indusia of narrow, filamentous segments, these uniseriate for most of length, composed of ± isodiametric cells, usually surpassing mature sporangia. Spores averaging 44–52 µm. 2n = 152.


Phenology: Sporulating summer–fall.
Habitat: Cliffs and rocky slopes, usually on sandstone or igneous substrates
Elevation: 300–3500 m

Distribution

V2 245-distribution-map.gif

Ariz., Colo., N.Mex., Okla., S.Dak., Tex.

Discussion

Woodsia neomexicana traditionally has been identified as W. mexicana. Both taxa are tetraploid and may share one parent (M. D. Windham 1993); W. neomexicana is separated from typical W. mexicana by its completely filamentous indusial segments, reduced glandularity, and more northerly distribution. Isozyme data suggest that W. neomexicana is an allotetraploid hybrid between W. phillipsii and the diploid progenitor of W. oregana subsp. cathcartiana (M. D. Windham 1993). As with all allopolyploids, W. neomexicana can vary in the direction of either parent, and some plants (especially those resembling W. phillipsii) can be difficult to identify. All characters except those controlled directly by ploidy level show this tendency, and spore size remains the most dependable character for distinguishing W. phillipsii and W. neomexicana. This species hybridizes with W. oregana subsp. cathcartiana and W. phillipsii to produce sterile tetraploids and triploids, respectively.

Selected References

None.

Lower Taxa

None.