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| − | --><span class="statement" id="st-d0_s0" data-properties="stem architecture;stem coloration;stem shape;stem shape;stem shape;stem shape;stem shape;stem atypical length;stem length;stem atypical width;stem width;stem pubescence"><b>Stems </b>usually unbranched, green, depressed-spheric or elliptic to shortly cylindric, ovoid when young, 2.9–8.5 (–10.4) × 1.8–7.5 (–8) cm, somewhat glaucous;</span> <span class="statement" id="st-d0_s1" data-properties="rib count;rib prominence;rib development;tubercle prominence">ribs usually 13, obscure to well developed, tubercles evident on ribs.</span> <span class="statement" id="st-d0_s2" data-properties=""><b>Spines </b>not obscuring stems;</span> <span class="statement" id="st-d0_s3" data-properties="spine arrangement;spine atypical count;spine atypical count;spine count;spine shape;spine shape;spine shape">radial spines (5–) 6–7 (–13) per areole, acicular, elliptic or rhombic in cross-section;</span> <span class="statement" id="st-d0_s4" data-properties="central spine atypical count;central spine atypical count;central spine count;abaxial central spine count;abaxial central spine shape;abaxial central spine course;abaxial central spine orientation;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine coloration;abaxial central spine shape;abaxial central spine orientation;abaxial central spine shape;abaxial central spine shape;abaxial central spine atypical length;abaxial central spine length;abaxial central spine width">central spines (0–) 1 (–3) per areole, abaxial central spine, if present, hooked or curving somewhat downward, straw colored to brown, highlighted purplish or reddish, hooked, porrect, terete or somewhat angled, 3–31 (–39) × 0.5–1 mm;</span> <span class="statement" id="st-d0_s5" data-properties="lateral central spine count;lateral central spine orientation;lateral central spine shape;lateral central spine prominence;lateral central spine shape;lateral central spine some measurement;abaxial spine course;cross-section count">lateral central spines 0–2 per areole, erect, similar to abaxial but not curved, angled to flat, somewhat inconspicuous, triangular in cross-section, 15–35 1–1.5 mm;</span> <span class="statement" id="st-d0_s6" data-properties="adaxial central spine variability">adaxial central spines not differentiated.</span> <span class="statement" id="st-d0_s7" data-properties="flower shape;flower atypical length;flower length;flower atypical width;flower width"><b>Flowers </b>campanulate, 2–3 (–4) × 1.1–3 (–3.5) cm;</span> <span class="statement" id="st-d0_s8" data-properties="outer tepal coloration;outer tepal coloration;outer tepal coloration;outer tepal coloration;outer tepal coloration;outer tepal coloration;outer tepal shape;outer tepal length;outer tepal atypical width;outer tepal width;outer tepal shape;outer tepal texture;outer tepal shape;outer tepal shape;midstripe coloration">outer tepals greenish to purple with brownish midstripes, pink, purple or whitish at margins, oblanceolate, 7–16 × 2–6 (–7) mm, mucronate, marginally membranous and crisped or minutely toothed;</span> <span class="statement" id="st-d0_s9" data-properties="inner tepal coloration or density;inner tepal coloration;largest tepal shape;largest tepal atypical length;largest tepal length;largest tepal width;largest tepal shape;largest tepal shape">inner tepals purple, sometimes suffused with brown, largest tepals oblanceolate, 10–22 (–30) × 4–7 mm, mucronate, somewhat irregularly toothed;</span> <span class="statement" id="st-d0_s10" data-properties="filament coloration;filament coloration;filament coloration;filament coloration;filament coloration">filaments white, tinged with pink to pink-purple;</span> <span class="statement" id="st-d0_s11" data-properties="anther coloration">anthers yellow.</span> <span class="statement" id="st-d0_s12" data-properties="fruit dehiscence;fruit coloration;fruit coloration;fruit coloration;fruit coloration;fruit shape;fruit length;fruit width;fruit condition or texture"><b>Fruits </b>irregularly dehiscent, green to tan, sometimes suffused with pink, shortly cylindric, 7–15 × 5–12 mm, dry;</span> <span class="statement" id="st-d0_s13" data-properties="scale count;scale texture;scale architecture">scales few, membranous, scarious-margined.</span> <span class="statement" id="st-d0_s14" data-properties="seed atypical length;seed length;seed atypical width;seed width"><b>Seeds </b>(1.2–) 1.8–2.7 × (1.9–) 2.5–3.8 mm;</span> <span class="statement" id="st-d0_s15" data-properties="cell shape;cell shape">cells convex but flattened apically.</span><!-- | + | --><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>usually unbranched, green, depressed-spheric or elliptic to shortly cylindric, ovoid when young, 2.9–8.5(–10.4) × 1.8–7.5(–8) cm, somewhat glaucous; ribs usually 13, obscure to well developed, tubercles evident on ribs. <b>Spines</b> not obscuring stems; radial spines (5–)6–7(–13) per areole, acicular, elliptic or rhombic in cross section; central spines (0–)1(–3) per areole, abaxial central spine, if present, hooked or curving somewhat downward, straw colored to brown, highlighted purplish or reddish, hooked, porrect, terete or somewhat angled, 3–31(–39) × 0.5–1 mm; lateral central spines 0–2 per areole, erect, similar to abaxial but not curved, angled to flat, somewhat inconspicuous, triangular in cross section, 15–35 1–1.5 mm; adaxial central spines not differentiated. <b>Flowers</b> campanulate, 2–3(–4) × 1.1–3(–3.5) cm; outer tepals greenish to purple with brownish midstripes, pink, purple or whitish at margins, oblanceolate, 7–16 × 2–6(–7) mm, mucronate, marginally membranous and crisped or minutely toothed; inner tepals purple, sometimes suffused with brown, largest tepals oblanceolate, 10–22(–30) × 4–7 mm, mucronate, somewhat irregularly toothed; filaments white, tinged with pink to pink-purple; anthers yellow. <b>Fruits</b> irregularly dehiscent, green to tan, sometimes suffused with pink, shortly cylindric, 7–15 × 5–12 mm, dry; scales few, membranous, scarious-margined. <b>Seeds</b> (1.2–)1.8–2.7 × (1.9–)2.5–3.8 mm; cells convex but flattened apically.</span><!-- |
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| | -->{{Treatment/Body | | -->{{Treatment/Body |
| | + | |phenology=Flowering late Apr–May. |
| | |habitat=Exposed clay hills, saltbush or sagebrush flats | | |habitat=Exposed clay hills, saltbush or sagebrush flats |
| | |elevation=1400-1500 m | | |elevation=1400-1500 m |
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| | |basionyms=Sclerocactus wetlandicus var. ilseae | | |basionyms=Sclerocactus wetlandicus var. ilseae |
| | |family=Cactaceae | | |family=Cactaceae |
| | + | |phenology=Flowering late Apr–May. |
| | |habitat=Exposed clay hills, saltbush or sagebrush flats | | |habitat=Exposed clay hills, saltbush or sagebrush flats |
| | |elevation=1400-1500 m | | |elevation=1400-1500 m |
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| | |publication year=1994 | | |publication year=1994 |
| | |special status= | | |special status= |
| − | |source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_375.xml | + | |source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V4/V4_375.xml |
| | |subfamily=Cactaceae subfam. Cactoideae | | |subfamily=Cactaceae subfam. Cactoideae |
| | |genus=Sclerocactus | | |genus=Sclerocactus |
| | |species=Sclerocactus brevispinus | | |species=Sclerocactus brevispinus |
| − | |abaxial central spine atypical length=31mm;39mm
| |
| − | |abaxial central spine coloration=straw colored;brown highlighted purplish or reddish
| |
| − | |abaxial central spine count=present
| |
| − | |abaxial central spine course=curving
| |
| − | |abaxial central spine length=3mm;31mm
| |
| − | |abaxial central spine orientation=porrect;downward
| |
| − | |abaxial central spine shape=angled;terete;hooked;hooked
| |
| − | |abaxial central spine width=0.5mm;1mm
| |
| − | |abaxial spine course=not curved
| |
| − | |adaxial central spine variability=not differentiated
| |
| − | |anther coloration=yellow
| |
| − | |cell shape=flattened;convex
| |
| − | |central spine atypical count=1;3
| |
| − | |central spine count=1
| |
| − | |cross-section count=15;35
| |
| − | |filament coloration=tinged with pink;pink-purple
| |
| − | |flower atypical length=3cm;4cm
| |
| − | |flower atypical width=3cm;3.5cm
| |
| − | |flower length=2cm;3cm
| |
| − | |flower shape=campanulate
| |
| − | |flower width=1.1cm;3cm
| |
| − | |fruit coloration=suffused with pink;green;tan
| |
| − | |fruit condition or texture=dry
| |
| − | |fruit dehiscence=dehiscent
| |
| − | |fruit length=7mm;15mm
| |
| − | |fruit shape=cylindric
| |
| − | |fruit width=5mm;12mm
| |
| − | |inner tepal coloration=suffused with brown
| |
| − | |inner tepal coloration or density=purple
| |
| − | |largest tepal atypical length=22mm;30mm
| |
| − | |largest tepal length=10mm;22mm
| |
| − | |largest tepal shape=toothed;mucronate;oblanceolate
| |
| − | |largest tepal width=4mm;7mm
| |
| − | |lateral central spine count=0;2
| |
| − | |lateral central spine orientation=erect
| |
| − | |lateral central spine prominence=inconspicuous
| |
| − | |lateral central spine shape=triangular;angled to flat
| |
| − | |lateral central spine some measurement=1mm;1.5mm
| |
| − | |midstripe coloration=brownish
| |
| − | |outer tepal atypical width=6mm;7mm
| |
| − | |outer tepal coloration=whitish;purple;pink;greenish;purple
| |
| − | |outer tepal length=7mm;16mm
| |
| − | |outer tepal shape=toothed;crisped;mucronate;oblanceolate
| |
| − | |outer tepal texture=membranous
| |
| − | |outer tepal width=2mm;6mm
| |
| − | |rib count=13
| |
| − | |rib development=developed
| |
| − | |rib prominence=obscure
| |
| − | |scale architecture=scarious-margined
| |
| − | |scale count=few
| |
| − | |scale texture=membranous
| |
| − | |seed atypical length=1.2mm;1.8mm
| |
| − | |seed atypical width=1.9mm;2.5mm
| |
| − | |seed length=1.8mm;2.7mm
| |
| − | |seed width=2.5mm;3.8mm
| |
| − | |spine arrangement=radial
| |
| − | |spine atypical count=7;13
| |
| − | |spine count=6;7
| |
| − | |spine shape=rhombic;elliptic;acicular
| |
| − | |stem architecture=unbranched
| |
| − | |stem atypical length=8.5cm;10.4cm
| |
| − | |stem atypical width=7.5cm;8cm
| |
| − | |stem coloration=green
| |
| − | |stem length=2.9cm;8.5cm
| |
| − | |stem pubescence=glaucous
| |
| − | |stem shape=ovoid;elliptic;shortly cylindric
| |
| − | |stem width=1.8cm;7.5cm
| |
| − | |tubercle prominence=evident
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| | -->[[Category:Treatment]][[Category:Sclerocactus]] | | -->[[Category:Treatment]][[Category:Sclerocactus]] |
Stems usually unbranched, green, depressed-spheric or elliptic to shortly cylindric, ovoid when young, 2.9–8.5(–10.4) × 1.8–7.5(–8) cm, somewhat glaucous; ribs usually 13, obscure to well developed, tubercles evident on ribs. Spines not obscuring stems; radial spines (5–)6–7(–13) per areole, acicular, elliptic or rhombic in cross section; central spines (0–)1(–3) per areole, abaxial central spine, if present, hooked or curving somewhat downward, straw colored to brown, highlighted purplish or reddish, hooked, porrect, terete or somewhat angled, 3–31(–39) × 0.5–1 mm; lateral central spines 0–2 per areole, erect, similar to abaxial but not curved, angled to flat, somewhat inconspicuous, triangular in cross section, 15–35 1–1.5 mm; adaxial central spines not differentiated. Flowers campanulate, 2–3(–4) × 1.1–3(–3.5) cm; outer tepals greenish to purple with brownish midstripes, pink, purple or whitish at margins, oblanceolate, 7–16 × 2–6(–7) mm, mucronate, marginally membranous and crisped or minutely toothed; inner tepals purple, sometimes suffused with brown, largest tepals oblanceolate, 10–22(–30) × 4–7 mm, mucronate, somewhat irregularly toothed; filaments white, tinged with pink to pink-purple; anthers yellow. Fruits irregularly dehiscent, green to tan, sometimes suffused with pink, shortly cylindric, 7–15 × 5–12 mm, dry; scales few, membranous, scarious-margined. Seeds (1.2–)1.8–2.7 × (1.9–)2.5–3.8 mm; cells convex but flattened apically.
Phenology: Flowering late Apr–May.
Habitat: Exposed clay hills, saltbush or sagebrush flats
Elevation: 1400-1500 m
Discussion
Of conservation concern.
The combination of spheric stems, short spines, and small pink flowers make Sclerocactus brevispinus easily identified. Morphologically, S. brevispinus is very similar to S. mesae-verdae. This similarity appears to be a case of convergent evolution. Phylogenetic analyses of chloroplast DNA sequences (J. M. Porter et al. 2000) support more recent common ancestry of S. brevispinus with S. whipplei, S. cloverae, S. glaucus, S. parviflorus, S. wetlandicus, and S. wrightiae than with S. mesae-verdae.
There is a well-documented cline of genetic mixing between Sclerocactus brevispinus and S. wetlandicus. An important natural threat to S. brevispinus is genetic swamping from the more widespread S. wetlandicus.
Sclerocactus brevispinus is considered to be conspecific with S. glaucus by the U.S. Fish and Wildlife Service, and is therefore protected with that threatened species.
Selected References
None.
