Difference between revisions of "Pachycereus schottii"

(Engelmann) D. R. Hunt

Bradleya 5: 93. 1987.

Common names: Senita old-man-cactus
Basionym: Cereus schottii Engelmann
Synonyms: Lophocereus schottii (Engelmann) Britton & RosePilocereuss chottii (Engelmann) Lemaire
Treatment appears in FNA Volume 4. Treatment on page 184. Mentioned on page 181, 183.
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--><span class="statement" id="st-d0_s0" data-properties="basal branch architecture"><b>Shrubs </b>[to small trees], with basal branches sometimes rooting to form thickets.</span> <span class="statement" id="st-d0_s1" data-properties="stem coloration;stem life cycle;stem coloration;areole life cycle;chevron prominence;chevron texture"><b>Stems </b>green with red around young areoles, aging gray-green with faint waxy chevrons;</span> <span class="statement" id="st-d0_s2" data-properties="rib shape;margin prominence or shape">ribs somewhat rounded, margins relatively flat;</span> <span class="statement" id="st-d0_s3" data-properties="areole some measurement;areole arrangement;areole shape;areole arrangement or shape;proximal stem width;distal stem arrangement">areoles 1–1.5 cm apart on rib crest, shield-shaped, on proximal stems 1 cm wide, those on distal stems closer together, circular;</span> <span class="statement" id="st-d0_s4" data-properties="hair life cycle;hair coloration;hair height or length or size">young hairs whitish gray, short.</span> <span class="statement" id="st-d0_s5" data-properties="spine growth form;spine coloration;spine life cycle;spine coloration;spine fragility or size;spine some measurement;spine some measurement;spine variability;spine coloration;spine coloration;spine coloration;spine width;spine shape;spine architecture;spine some measurement;proximal stem count;central spine arrangement;distal stem count"><b>Spines </b>strongly dimorphic, those on proximal stems usually 7–10 per areole, rose-red, aging gray, stout, usually 5–7 mm, to less than 30 mm, weakly differentiated as central and radial spines, those on distal stems usually more than 15 per areole, gray to blackish (rarely aging copper-brown), thin, flattened, twisted, to 7.5 cm;</span> <span class="statement" id="st-d0_s6" data-properties="bristle some measurement">bristles 4–10 cm.</span> <span class="statement" id="st-d0_s7" data-properties="flower position;flower count;flower height or length or size;flower shape;flower length;flower width;areole development;areole duration"><b>Flowers </b>lateral on stems, often several per areole, on specialized bristle-forming or transition areoles, short funnelform, 3–4.5 × 2–3 cm;</span> <span class="statement" id="st-d0_s8" data-properties="flower tube width;flower tube architecture;flower tube size or width;flower tube some measurement">flower tube thin walled, very narrow, less than 1.5 cm;</span> <span class="statement" id="st-d0_s9" data-properties="nectar chamber architecture">nectar chamber open;</span> <span class="statement" id="st-d0_s10" data-properties="margin coloration">scales rose with green margins;</span> <span class="statement" id="st-d0_s11" data-properties="tepal orientation;tepal orientation">tepals spreading, reflexed by morning;</span> <span class="statement" id="st-d0_s12" data-properties="outer tepal shape;outer tepal relief">outer tepals somewhat fimbriate, minutely papillate;</span> <span class="statement" id="st-d0_s13" data-properties="inner tepal count;inner tepal some measurement;center coloration;apex shape">inner tepals 25, with pink-orange center, 11–16 mm, apex acute;</span> <span class="statement" id="st-d0_s14" data-properties="stamen count;stamen coloration;stamen coloration;stamen some measurement">stamens 100, white, turning pink by next morning, to 2 cm;</span> <span class="statement" id="st-d0_s15" data-properties="anther coloration">anthers cream;</span> <span class="statement" id="st-d0_s16" data-properties="ovary size;tubercle count;spine count">ovary small, tubercles and spines absent;</span> <span class="statement" id="st-d0_s17" data-properties="style coloration;style height or length or size;style size or width">styles white, short, narrow;</span> <span class="statement" id="st-d0_s18" data-properties="stigma lobe orientation;stigma lobe height or length or size">stigma lobes erect, relatively short.</span> <span class="statement" id="st-d0_s19" data-properties="fruit coloration;fruit coloration;fruit shape;fruit shape;fruit shape;fruit some measurement"><b>Fruits </b>candy-apple red, when immature greenish, ovoid to subspheric, 25–40 mm.</span> <span class="statement" id="st-d0_s20" data-properties="raphe prominence"><b>Seeds </b>helmet-shaped, often with a prominent raphe.</span> <span class="statement" id="st-d0_s21" data-properties="seed shape;2n chromosome count">2n = 22.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Shrubs </b>[to small trees], with basal branches sometimes rooting to form thickets. <b>Stems</b> green with red around young areoles, aging gray-green with faint waxy chevrons; ribs somewhat rounded, margins relatively flat; areoles 1–1.5 cm apart on rib crest, shield-shaped, on proximal stems 1 cm wide, those on distal stems closer together, circular; young hairs whitish gray, short. <b>Spines</b> strongly dimorphic, those on proximal stems usually 7–10 per areole, rose-red, aging gray, stout, usually 5–7 mm, to less than 30 mm, weakly differentiated as central and radial spines, those on distal stems usually more than 15 per areole, gray to blackish (rarely aging copper-brown), thin, flattened, twisted, to 7.5 cm; bristles 4–10 cm. <b>Flowers</b> lateral on stems, often several per areole, on specialized bristle-forming or transition areoles, short funnelform, 3–4.5 × 2–3 cm; flower tube thin walled, very narrow, less than 1.5 cm; nectar chamber open; scales rose with green margins; tepals spreading, reflexed by morning; outer tepals somewhat fimbriate, minutely papillate; inner tepals 25, with pink-orange center, 11–16 mm, apex acute; stamens 100, white, turning pink by next morning, to 2 cm; anthers cream; ovary small, tubercles and spines absent; styles white, short, narrow; stigma lobes erect, relatively short. <b>Fruits</b> candy-apple red, when immature greenish, ovoid to subspheric, 25–40 mm. <b>Seeds</b> helmet-shaped, often with a prominent raphe. <b>2n</b> = 22.</span><!--
  
 
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|phenology=Flowering late Mar–Sep[-Dec]; fruiting often 1 month after flowering.
 
|habitat=Cercidium-Larrea-Olneya-Carnegiea communities, around washes, on sandy [gravelly] soils
 
|habitat=Cercidium-Larrea-Olneya-Carnegiea communities, around washes, on sandy [gravelly] soils
 
|elevation=400-500 m
 
|elevation=400-500 m
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|basionyms=Cereus schottii
 
|basionyms=Cereus schottii
 
|family=Cactaceae
 
|family=Cactaceae
 +
|phenology=Flowering late Mar–Sep[-Dec]; fruiting often 1 month after flowering.
 
|habitat=Cercidium-Larrea-Olneya-Carnegiea communities, around washes, on sandy [gravelly] soils
 
|habitat=Cercidium-Larrea-Olneya-Carnegiea communities, around washes, on sandy [gravelly] soils
 
|elevation=400-500 m
 
|elevation=400-500 m
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|publication year=1987
 
|publication year=1987
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_348.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V4/V4_348.xml
 
|subfamily=Cactaceae subfam. Cactoideae
 
|subfamily=Cactaceae subfam. Cactoideae
 
|genus=Pachycereus
 
|genus=Pachycereus
 
|species=Pachycereus schottii
 
|species=Pachycereus schottii
|2n chromosome count=22
 
|anther coloration=cream
 
|apex shape=acute
 
|areole arrangement=apart
 
|areole arrangement or shape=circular
 
|areole development=specialized
 
|areole duration=transition
 
|areole life cycle=young
 
|areole shape=shield-shaped
 
|areole some measurement=1cm;1.5cm
 
|basal branch architecture=rooting
 
|bristle some measurement=4cm;10cm
 
|center coloration=pink-orange
 
|central spine arrangement=radial
 
|chevron prominence=faint
 
|chevron texture=waxy
 
|distal stem arrangement=closer
 
|flower count=several
 
|flower height or length or size=short
 
|flower length=3cm;4.5cm
 
|flower position=lateral
 
|flower shape=funnelform
 
|flower tube architecture=walled
 
|flower tube size or width=narrow
 
|flower tube some measurement=0cm;1.5cm
 
|flower tube width=thin
 
|flower width=2cm;3cm
 
|fruit coloration=greenish;candy-apple red
 
|fruit shape=ovoid;subspheric
 
|fruit some measurement=25mm;40mm
 
|hair coloration=whitish gray
 
|hair height or length or size=short
 
|hair life cycle=young
 
|inner tepal count=25
 
|inner tepal some measurement=11mm;16mm
 
|margin coloration=green
 
|margin prominence or shape=flat
 
|nectar chamber architecture=open
 
|outer tepal relief=papillate
 
|outer tepal shape=fimbriate
 
|ovary size=small
 
|proximal stem count=7;10
 
|proximal stem width=1
 
|raphe prominence=prominent
 
|rib shape=rounded
 
|seed shape=helmet-shaped
 
|spine architecture=twisted
 
|spine coloration=gray;blackish
 
|spine count=absent
 
|spine fragility or size=stout
 
|spine growth form=dimorphic
 
|spine life cycle=aging
 
|spine shape=flattened
 
|spine some measurement=0cm;7.5cm
 
|spine variability=differentiated
 
|spine width=thin
 
|stamen coloration=pink;white
 
|stamen count=100
 
|stamen some measurement=0cm;2cm
 
|stem coloration=gray-green;green
 
|stem life cycle=aging
 
|stigma lobe height or length or size=short
 
|stigma lobe orientation=erect
 
|style coloration=white
 
|style height or length or size=short
 
|style size or width=narrow
 
|tepal orientation=reflexed;spreading
 
|tubercle count=absent
 
 
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Revision as of 14:06, 27 July 2019

Shrubs [to small trees], with basal branches sometimes rooting to form thickets. Stems green with red around young areoles, aging gray-green with faint waxy chevrons; ribs somewhat rounded, margins relatively flat; areoles 1–1.5 cm apart on rib crest, shield-shaped, on proximal stems 1 cm wide, those on distal stems closer together, circular; young hairs whitish gray, short. Spines strongly dimorphic, those on proximal stems usually 7–10 per areole, rose-red, aging gray, stout, usually 5–7 mm, to less than 30 mm, weakly differentiated as central and radial spines, those on distal stems usually more than 15 per areole, gray to blackish (rarely aging copper-brown), thin, flattened, twisted, to 7.5 cm; bristles 4–10 cm. Flowers lateral on stems, often several per areole, on specialized bristle-forming or transition areoles, short funnelform, 3–4.5 × 2–3 cm; flower tube thin walled, very narrow, less than 1.5 cm; nectar chamber open; scales rose with green margins; tepals spreading, reflexed by morning; outer tepals somewhat fimbriate, minutely papillate; inner tepals 25, with pink-orange center, 11–16 mm, apex acute; stamens 100, white, turning pink by next morning, to 2 cm; anthers cream; ovary small, tubercles and spines absent; styles white, short, narrow; stigma lobes erect, relatively short. Fruits candy-apple red, when immature greenish, ovoid to subspheric, 25–40 mm. Seeds helmet-shaped, often with a prominent raphe. 2n = 22.


Phenology: Flowering late Mar–Sep[-Dec]; fruiting often 1 month after flowering.
Habitat: Cercidium-Larrea-Olneya-Carnegiea communities, around washes, on sandy [gravelly] soils
Elevation: 400-500 m

Distribution

V4 348-distribution-map.gif

Ariz., Mexico (Baja California, Baja California Sur, Sinaloa, Sonora).

Discussion

Pachycereus schottii is primarily a Mexican species occuring in southernmost Arizona only in Organ Pipe Cactus National Monument at nine sites very close to the international boundary, where winter temperatures do not generally reach freezing. The Arizona populations do not appear to be experiencing much recruitment via seeds, but persist by vegetative propagation. In Mexico, plant height, rib number, and rib shape vary considerably with microhabitat. In subtropical thorn forest P. schottii may be a small tree with a very short trunk. Three varieties of P. schottii were formally recognized (G. S. Lindsay 1963) but need to be re-examined using DNA techniques. If varieties or subspecies are accepted, Arizona populations would represent the typical form.

In contrast to other species of Pachycereus, P. schottii and its narrowly restricted cousin P. gatesii of Baja California Sur, Mexico have the smallest flowers and seeds and the most fleshy and naked fruits. The flowering portion of each adult stem always has long, grayish bristles, inspiring the anthropomorphic vernacular names referring to old age.

The moth Upiga virescens (Pyralidae) is the obligate pollinator of Pachycereus schottii, forming a mutualistic system comparable with the yucca and yucca moth (J. N. Holland and T. H. Fleming 1999, 1999b; R. S. Folger and A. D. Zimmerman 2000). In populations near the international border, flowers of P. schottii open ten minutes after sunset (J. N. Holland, pers. comm.).

Selected References

None.

Lower Taxa

None.
... more about "Pachycereus schottii"
Arthur C. Gibson +
(Engelmann) D. R. Hunt +
Cereus schottii +
Senita +  and old-man-cactus +
Ariz. +, Mexico (Baja California +, Baja California Sur +, Sinaloa +  and Sonora). +
400-500 m +
Cercidium-Larrea-Olneya-Carnegiea communities, around washes, on sandy [gravelly] soils +
Flowering late Mar–Sep[-Dec] +  and fruiting often 1 month after flowering. +
Lophocereus schottii +  and Pilocereuss chottii +
Pachycereus schottii +
Pachycereus +
species +