Difference between revisions of "Fumariaceae"

Linnaeus
Common names: Fumitory Family
Treatment appears in FNA Volume 3. Treatment on page 340.
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--><span class="statement" id="st-d0_s0" data-properties="herb duration;herb duration;herb architecture;herb architecture"><b>Herbs,</b> annual or perennial, scapose or caulescent, from taproots, bulblets, tubers, or rhizomes;</span> <span class="statement" id="st-d0_s1" data-properties="sap coloration">sap clear.</span> <span class="statement" id="st-d0_s2" data-properties="stem architecture;stem orientation;stem growth form;stem growth form;stem architecture;stem architecture"><b>Stems </b>when present leafy, erect to prostrate or climbing, simple or branching.</span> <span class="statement" id="st-d0_s3" data-properties="leaf position;leaf position;leaf arrangement;leaf architecture;leaf architecture;leaf architecture"><b>Leaves </b>basal and/or cauline, alternate, mostly compound, sometimes simple, without stipules, petiolate;</span> <span class="statement" id="st-d0_s4" data-properties="order quantity;order architecture">blade with 2-6 odd-pinnate orders of leaflets and/or lobes.</span> <span class="statement" id="st-d0_s5" data-properties="inflorescence position;inflorescence position;inflorescence position;inflorescence position;inflorescence position;inflorescence position;inflorescence architecture;inflorescence architecture;inflorescence architecture;inflorescence architecture;inflorescence arrangement;inflorescence arrangement;inflorescence arrangement;inflorescence arrangement;inflorescence arrangement"><b>Inflorescences </b>terminal, axillary, extra-axillary, or leaf-opposed, unifloral or else multifloral and thyrsoid, paniculate, racemose, or corymbose;</span> <span class="statement" id="st-d0_s6" data-properties="peduncle presence">peduncles present;</span> <span class="statement" id="st-d0_s7" data-properties="beak presence">bracts present.</span> <span class="statement" id="st-d0_s8" data-properties="flower architecture or shape;flower quantity;flower shape;flower shape;plane quantity;plane orientation"><b>Flowers </b>bilaterally symmetric about 1 plane or each of 2 perpendicular planes;</span> <span class="statement" id="st-d0_s9" data-properties="pedicel presence">pedicel present;</span> <span class="statement" id="st-d0_s10" data-properties="sepal duration;sepal duration;sepal quantity;sepal width">sepals caducous or persistent, 2, thin;</span> <span class="statement" id="st-d0_s11" data-properties="petal quantity;petal fusion;petal fusion;petal fusion;whorl quantity;whorl quantity">petals 4, distinct or coherent basally to almost completely connate, in 2 whorls of 2;</span> <span class="statement" id="st-d0_s12" data-properties="outer petal variability;outer petal variability;outer petal quantity;outer petal quantity;outer petal shape;outer petal shape">outer petals alike or dissimilar, 1 or both sometimes swollen or spurred basally;</span> <span class="statement" id="st-d0_s13" data-properties="inner petal variability;inner petal fusion;inner petal shape;abaxial median crest architecture;abaxial median crest texture;abaxial median crest relief">inner petals alike, apically connate, clawed, with somewhat hollow, membranous, wrinkled, abaxial median crests;</span> <span class="statement" id="st-d0_s14" data-properties="stamen quantity;bundle quantity;outer petal quantity;outer petal arrangement">stamens 6, in 2 bundles of 3 each, opposite outer petals;</span> <span class="statement" id="st-d0_s15" data-properties="bundle filament fusion;bundle filament fusion">filaments of each bundle partially to completely connate, sometimes basally adnate to petals, with basal nectariferous tissue often in form of spur;</span> <span class="statement" id="st-d0_s16" data-properties="anther arrangement;median anther architecture or structure in adjective form;lateral anther architecture or structure in adjective form">anthers connivent, adhering to stigma, median anthers 2-locular, lateral anthers 1-locular;</span> <span class="statement" id="st-d0_s17" data-properties="pistil quantity;pistil architecture">pistil 1, 2-carpellate;</span> <span class="statement" id="st-d0_s18" data-properties="ovary architecture or structure in adjective form">ovary 1-locular;</span> <span class="statement" id="st-d0_s19" data-properties="placenta arrangement">placentae parietal;</span> <span class="statement" id="st-d0_s20" data-properties="style shape;style texture">style threadlike, rigid;</span> <span class="statement" id="st-d0_s21" data-properties="stigma quantity;stigma shape;lobe quantity;apical horn quantity;stigmatic surface shape">stigma 1, compressed, with 2 lobes or apical horns, and/or 2-8 papillar stigmatic surfaces.</span> <span class="statement" id="st-d0_s22" data-properties="fruit architecture;fruit dehiscence;fruit dehiscence;fruit arrangement or dehiscence"><b>Fruits </b>capsular, indehiscent or dehiscent and valvate.</span> <span class="statement" id="st-d0_s23" data-properties="seed quantity;seed size;elaiosome presence"><b>Seeds </b>1-many, small, elaiosome (oil-bearing appendage) often present.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Herbs,</b> annual or perennial, scapose or caulescent, from taproots, bulblets, tubers, or rhizomes; sap clear. <b>Stems</b> when present leafy, erect to prostrate or climbing, simple or branching. <b>Leaves</b> basal and/or cauline, alternate, mostly compound, sometimes simple, without stipules, petiolate; blade with 2-6 odd-pinnate orders of leaflets and/or lobes. <b>Inflorescences</b> terminal, axillary, extra-axillary, or leaf-opposed, unifloral or else multifloral and thyrsoid, paniculate, racemose, or corymbose; peduncles present; bracts present. <b>Flowers</b> bilaterally symmetric about 1 plane or each of 2 perpendicular planes; pedicel present; sepals caducous or persistent, 2, thin; petals 4, distinct or coherent basally to almost completely connate, in 2 whorls of 2; outer petals alike or dissimilar, 1 or both sometimes swollen or spurred basally; inner petals alike, apically connate, clawed, with somewhat hollow, membranous, wrinkled, abaxial median crests; stamens 6, in 2 bundles of 3 each, opposite outer petals; filaments of each bundle partially to completely connate, sometimes basally adnate to petals, with basal nectariferous tissue often in form of spur; anthers connivent, adhering to stigma, median anthers 2-locular, lateral anthers 1-locular; pistil 1, 2-carpellate; ovary 1-locular; placentae parietal; style threadlike, rigid; stigma 1, compressed, with 2 lobes or apical horns, and/or 2-8 papillar stigmatic surfaces. <b>Fruits</b> capsular, indehiscent or dehiscent and valvate. <b>Seeds</b> 1-many, small, elaiosome (oil-bearing appendage) often present.</span><!--
  
 
-->{{Treatment/Body
 
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|distribution=North America;Eurasia
+
|distribution=North America;Eurasia.
 
|discussion=<p>Genera 19, species ca. 450 (4 genera, 23 species in the flora).</p><!--
 
|discussion=<p>Genera 19, species ca. 450 (4 genera, 23 species in the flora).</p><!--
 
--><p>The genera of Fumariaceae are distributed mostly in the Old World and primarily in temperate Eurasia. One acaulescent species of Dicentra occurs in Siberia, Kamchatka, and Japan; a caulescent species is found in western China and northern Burma; and nine climbing species are distributed throughout the Himalayan area and Burma. More than 400 taxa of Corydalis and 50 of Fumaria, distributed primarily throughout temperate, often montane, regions of Eurasia and Africa, have been described. Adlumia comprises only two species, which are quite similar morphologically, one from North America and the other from East Asia.</p><!--
 
--><p>The genera of Fumariaceae are distributed mostly in the Old World and primarily in temperate Eurasia. One acaulescent species of Dicentra occurs in Siberia, Kamchatka, and Japan; a caulescent species is found in western China and northern Burma; and nine climbing species are distributed throughout the Himalayan area and Burma. More than 400 taxa of Corydalis and 50 of Fumaria, distributed primarily throughout temperate, often montane, regions of Eurasia and Africa, have been described. Adlumia comprises only two species, which are quite similar morphologically, one from North America and the other from East Asia.</p><!--
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|family=Fumariaceae
 
|family=Fumariaceae
 
|illustrator=John Myers
 
|illustrator=John Myers
|distribution=North America;Eurasia
+
|distribution=North America;Eurasia.
 
|reference=ernst1962a;fedde1936a;gunn1980a;hutchinson1921a;liden1986a;rachelle1974a;ryberg1960a;ryberg1960b
 
|reference=ernst1962a;fedde1936a;gunn1980a;hutchinson1921a;liden1986a;rachelle1974a;ryberg1960a;ryberg1960b
 
|publication title=
 
|publication title=
 
|publication year=
 
|publication year=
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V3/V3_145.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V3/V3_145.xml
|abaxial median crest architecture=hollow
 
|abaxial median crest relief=wrinkled
 
|abaxial median crest texture=membranous
 
|anther arrangement=connivent
 
|apical horn quantity=2;8
 
|beak presence=absent
 
|bundle filament fusion=adnate;connate
 
|bundle quantity=2
 
|elaiosome presence=absent
 
|flower architecture or shape=symmetric
 
|flower quantity=1
 
|flower shape=each;plane
 
|fruit architecture=capsular
 
|fruit arrangement or dehiscence=valvate
 
|fruit dehiscence=dehiscent;indehiscent
 
|herb architecture=caulescent;scapose
 
|herb duration=perennial;annual
 
|inflorescence architecture=thyrsoid;multifloral;else;unifloral
 
|inflorescence arrangement=corymbose;racemose;corymbose;racemose;paniculate
 
|inflorescence position=leaf-opposed;extra-axillary;leaf-opposed;extra-axillary;axillary;terminal
 
|inner petal fusion=connate
 
|inner petal shape=clawed
 
|inner petal variability=alike
 
|lateral anther architecture or structure in adjective form=1-locular
 
|leaf architecture=petiolate;simple;compound
 
|leaf arrangement=alternate
 
|leaf position=cauline;basal
 
|lobe quantity=2
 
|median anther architecture or structure in adjective form=2-locular
 
|order architecture=odd-pinnate
 
|order quantity=2;6
 
|outer petal arrangement=opposite
 
|outer petal quantity=3;both;1
 
|outer petal shape=spurred;swollen
 
|outer petal variability=dissimilar;alike
 
|ovary architecture or structure in adjective form=1-locular
 
|pedicel presence=absent
 
|peduncle presence=absent
 
|petal fusion=connate;coherent;distinct
 
|petal quantity=4
 
|pistil architecture=2-carpellate
 
|pistil quantity=1
 
|placenta arrangement=parietal
 
|plane orientation=perpendicular
 
|plane quantity=2
 
|sap coloration=clear
 
|seed quantity=1;many
 
|seed size=small
 
|sepal duration=persistent;caducous
 
|sepal quantity=2
 
|sepal width=thin
 
|stamen quantity=6
 
|stem architecture=branching;simple;leafy
 
|stem growth form=climbing;prostrate
 
|stem orientation=erect
 
|stigma quantity=1
 
|stigma shape=compressed
 
|stigmatic surface shape=papillar
 
|style shape=thread-like
 
|style texture=rigid
 
|whorl quantity=2;2
 
 
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Revision as of 13:34, 27 July 2019

Herbs, annual or perennial, scapose or caulescent, from taproots, bulblets, tubers, or rhizomes; sap clear. Stems when present leafy, erect to prostrate or climbing, simple or branching. Leaves basal and/or cauline, alternate, mostly compound, sometimes simple, without stipules, petiolate; blade with 2-6 odd-pinnate orders of leaflets and/or lobes. Inflorescences terminal, axillary, extra-axillary, or leaf-opposed, unifloral or else multifloral and thyrsoid, paniculate, racemose, or corymbose; peduncles present; bracts present. Flowers bilaterally symmetric about 1 plane or each of 2 perpendicular planes; pedicel present; sepals caducous or persistent, 2, thin; petals 4, distinct or coherent basally to almost completely connate, in 2 whorls of 2; outer petals alike or dissimilar, 1 or both sometimes swollen or spurred basally; inner petals alike, apically connate, clawed, with somewhat hollow, membranous, wrinkled, abaxial median crests; stamens 6, in 2 bundles of 3 each, opposite outer petals; filaments of each bundle partially to completely connate, sometimes basally adnate to petals, with basal nectariferous tissue often in form of spur; anthers connivent, adhering to stigma, median anthers 2-locular, lateral anthers 1-locular; pistil 1, 2-carpellate; ovary 1-locular; placentae parietal; style threadlike, rigid; stigma 1, compressed, with 2 lobes or apical horns, and/or 2-8 papillar stigmatic surfaces. Fruits capsular, indehiscent or dehiscent and valvate. Seeds 1-many, small, elaiosome (oil-bearing appendage) often present.

Distribution

North America, Eurasia.

Discussion

Genera 19, species ca. 450 (4 genera, 23 species in the flora).

The genera of Fumariaceae are distributed mostly in the Old World and primarily in temperate Eurasia. One acaulescent species of Dicentra occurs in Siberia, Kamchatka, and Japan; a caulescent species is found in western China and northern Burma; and nine climbing species are distributed throughout the Himalayan area and Burma. More than 400 taxa of Corydalis and 50 of Fumaria, distributed primarily throughout temperate, often montane, regions of Eurasia and Africa, have been described. Adlumia comprises only two species, which are quite similar morphologically, one from North America and the other from East Asia.

Most European and some American systematists treat Fumariaceae as a subfamily of Papaveraceae. However, although a few taxa are morphologically intermediate, the members of Fumariaceae generally are quite distinct from those of Papaveraceae in several respects, including floral symmetry, sap character, and stamen number and fusion.

Key

1 Petals almost completely connate, spongy; plants climbing, petiolules and reduced leaflets twining and tendril-like. Adlumia
1 Petals coherent or connate only basally, not spongy; plants not climbing. > 2
2 Both outer petals swollen or spurred basally. Dicentra
2 Only 1 outer petal swollen or spurred basally. > 3
3 Fruit an elongate, dehiscent capsule; seeds more than 1, with elaiosome. Corydalis
3 Fruit a ± globose, indehiscent capsule; seeds 1, without elaiosome. Fumaria