Difference between revisions of "Dryopteris carthusiana"

(Villars) H. P. Fuchs

Bull. Soc. Bot. France 105: 339. 1959.

Common names: Spinulose wood fern toothed wood fern dryoptère de cartheuser
Basionym: Polypodium carthusianum Villars Hist. Pl. Dauphiné 1: 292. 1786
Synonyms: Dryopteris austriaca var. spinulosa (O. F. Mueller) Fiori Dryopteris spinulosa (O. F. Mueller) Watt Polypodium spinulosum O. F. Mueller
Treatment appears in FNA Volume 2.
FNA>Volume Importer
 
m (Fixed Nfld. and Labr. distribution to match printed version.)
 
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}}
 
}}
 
|common_names=Spinulose wood fern;toothed wood fern;dryoptère de cartheuser
 
|common_names=Spinulose wood fern;toothed wood fern;dryoptère de cartheuser
|basionyms={{Treatment/ID/Synonym
+
|basionyms={{Treatment/ID/Basionym
 
|name=Polypodium carthusianum
 
|name=Polypodium carthusianum
 
|authority=Villars
 
|authority=Villars
 +
|rank=species
 +
|publication_title=Hist. Pl. Dauphiné
 +
|publication_place=1: 292. 1786
 
}}
 
}}
 
|synonyms={{Treatment/ID/Synonym
 
|synonyms={{Treatment/ID/Synonym
 
|name=Dryopteris austriaca var. spinulosa
 
|name=Dryopteris austriaca var. spinulosa
 
|authority=(O. F. Mueller) Fiori
 
|authority=(O. F. Mueller) Fiori
}}{{Treatment/ID/Synonym
+
|rank=variety
 +
}} {{Treatment/ID/Synonym
 
|name=Dryopteris spinulosa
 
|name=Dryopteris spinulosa
 
|authority=(O. F. Mueller) Watt
 
|authority=(O. F. Mueller) Watt
}}{{Treatment/ID/Synonym
+
|rank=species
 +
}} {{Treatment/ID/Synonym
 
|name=Polypodium spinulosum
 
|name=Polypodium spinulosum
 
|authority=O. F. Mueller
 
|authority=O. F. Mueller
 +
|rank=species
 
}}
 
}}
 
|hierarchy=Dryopteridaceae;Dryopteris;Dryopteris carthusiana
 
|hierarchy=Dryopteridaceae;Dryopteris;Dryopteris carthusiana
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}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="leaf architecture;leaf condition;leaf length;leaf width"><b>Leaves </b>monomorphic, dying in winter, 15–75 × 10–30 cm.</span> <span class="statement" id="st-d0_s1" data-properties="petiole length;petiole architecture or pubescence"><b>Petiole </b>1/4–1/3 length of leaf, scaly at least at base;</span> <span class="statement" id="st-d0_s2" data-properties="scale arrangement;scale coloration">scales scattered, tan.</span> <span class="statement" id="st-d0_s3" data-properties="blade coloration;blade shape;blade shape;blade growth form or texture;blade architecture or function or pubescence"><b>Blade </b>light green, ovatelanceolate, 2–3-pinnate-pinnatifid, herbaceous, not glandular.</span> <span class="statement" id="st-d0_s4" data-properties="pinna shape"><b>Pinnae </b>± in plane of blade, lance-oblong;</span> <span class="statement" id="st-d0_s5" data-properties="basal pinna shape;basal pinna size;basal pinnule length or size;pinnule arrangement;pinnule orientation;pinnule length or size;basal pinnule orientation">basal pinnae lanceolate-deltate, slightly reduced, basal pinnules usually longer than adjacent pinnules, basal basiscopic pinnule longer than basal acroscopic pinnule;</span> <span class="statement" id="st-d0_s6" data-properties="pinnule margin architecture or shape;tooth architecture or shape">pinnule margins serrate, teeth spiny.</span> <span class="statement" id="st-d0_s7" data-properties="sorus position"><b>Sori </b>midway between midvein and margin of segments.</span> <span class="statement" id="st-d0_s8" data-properties="gland count"><b>Indusia </b>lacking glands.</span> <span class="statement" id="st-d0_s9" data-properties="2n chromosome count">2n = 164.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Leaves </b>monomorphic, dying in winter, 15–75 × 10–30 cm. <b>Petiole</b> 1/4–1/3 length of leaf, scaly at least at base; scales scattered, tan. <b>Blade</b> light green, ovate-lanceolate, 2–3-pinnate-pinnatifid, herbaceous, not glandular. <b>Pinnae</b> ± in plane of blade, lance-oblong; basal pinnae lanceolate-deltate, slightly reduced, basal pinnules usually longer than adjacent pinnules, basal basiscopic pinnule longer than basal acroscopic pinnule; pinnule margins serrate, teeth spiny. <b>Sori</b> midway between midvein and margin of segments. <b>Indusia</b> lacking glands. <b>2n</b> = 164.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
 
|habitat=Swampy woods, moist wooded slopes, stream banks, and conifer plantations
 
|habitat=Swampy woods, moist wooded slopes, stream banks, and conifer plantations
 
|elevation=0–1200 m
 
|elevation=0–1200 m
|distribution=Alta.;B.C.;Man.;N.B.;Nfld.;N.W.T.;N.S.;Ont.;P.E.I.;Que.;Sask.;Yukon;Ark.;Conn.;Del.;Idaho;Ill.;Ind.;Iowa;Ky.;Maine;Md.;Mass.;Mich.;Minn.;Mo.;Mont.;Nebr.;N.H.;N.J.;N.Y.;N.C.;N.Dak.;Ohio;Pa.;R.I.;S.C.;Tenn.;Vt.;Va.;Wash.;W.Va.;Wis.;Eurasia.
+
|distribution=Alta.;B.C.;Man.;N.B.;Nfld. and Labr.;N.W.T.;N.S.;Ont.;P.E.I.;Que.;Sask.;Yukon;Ark.;Conn.;Del.;Idaho;Ill.;Ind.;Iowa;Ky.;Maine;Md.;Mass.;Mich.;Minn.;Mo.;Mont.;Nebr.;N.H.;N.J.;N.Y.;N.C.;N.Dak.;Ohio;Pa.;R.I.;S.C.;Tenn.;Vt.;Va.;Wash.;W.Va.;Wis.;Eurasia.
|discussion=<p>Dryopteris carthusiana is tetraploid. Dryopteris intermedia is one parent, as indicated by chromosome pairing in their hybrid D. × triploidea Wherry. The other parent is the hypothetical missing ancestral species " D. semicristata " (see discussion for D. cristata). Dryopteris carthusiana hybridizes with five species; hybrids can be separated from D. intermedia by the lack of glandular hairs and by having 2-pinnate leaves.</p>
+
|discussion=<p><i>Dryopteris carthusiana</i> is tetraploid. <i>Dryopteris intermedia</i> is one parent, as indicated by chromosome pairing in their hybrid D. × triploidea Wherry. The other parent is the hypothetical missing ancestral species " D. semicristata " (see discussion for <i>D. cristata</i>). <i>Dryopteris carthusiana</i> hybridizes with five species; hybrids can be separated from <i>D. intermedia</i> by the lack of glandular hairs and by having 2-pinnate leaves.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Dryopteris carthusiana
 
name=Dryopteris carthusiana
|author=
 
 
|authority=(Villars) H. P. Fuchs
 
|authority=(Villars) H. P. Fuchs
 
|rank=species
 
|rank=species
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|habitat=Swampy woods, moist wooded slopes, stream banks, and conifer plantations
 
|habitat=Swampy woods, moist wooded slopes, stream banks, and conifer plantations
 
|elevation=0–1200 m
 
|elevation=0–1200 m
|distribution=Alta.;B.C.;Man.;N.B.;Nfld.;N.W.T.;N.S.;Ont.;P.E.I.;Que.;Sask.;Yukon;Ark.;Conn.;Del.;Idaho;Ill.;Ind.;Iowa;Ky.;Maine;Md.;Mass.;Mich.;Minn.;Mo.;Mont.;Nebr.;N.H.;N.J.;N.Y.;N.C.;N.Dak.;Ohio;Pa.;R.I.;S.C.;Tenn.;Vt.;Va.;Wash.;W.Va.;Wis.;Eurasia.
+
|distribution=Alta.;B.C.;Man.;N.B.;Nfld. and Labr.;N.W.T.;N.S.;Ont.;P.E.I.;Que.;Sask.;Yukon;Ark.;Conn.;Del.;Idaho;Ill.;Ind.;Iowa;Ky.;Maine;Md.;Mass.;Mich.;Minn.;Mo.;Mont.;Nebr.;N.H.;N.J.;N.Y.;N.C.;N.Dak.;Ohio;Pa.;R.I.;S.C.;Tenn.;Vt.;Va.;Wash.;W.Va.;Wis.;Eurasia.
 
|reference=None
 
|reference=None
 
|publication title=Bull. Soc. Bot. France
 
|publication title=Bull. Soc. Bot. France
 
|publication year=1959
 
|publication year=1959
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_507.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_507.xml
 
|genus=Dryopteris
 
|genus=Dryopteris
 
|species=Dryopteris carthusiana
 
|species=Dryopteris carthusiana
|2n chromosome count=164
 
|basal pinna shape=lanceolate-deltate
 
|basal pinna size=reduced
 
|basal pinnule length or size=usually longer
 
|basal pinnule orientation=acroscopic
 
|blade architecture or function or pubescence=not glandular
 
|blade coloration=light green
 
|blade growth form or texture=herbaceous
 
|blade shape=2-3-pinnate-pinnatifid;ovatelanceolate
 
|gland count=lacking
 
|leaf architecture=monomorphic
 
|leaf condition=dying
 
|leaf length=15cm;75cm
 
|leaf width=10cm;30cm
 
|petiole architecture or pubescence=scaly
 
|petiole length=1/4 length of leaf;1/3 length of leaf
 
|pinna shape=lance-oblong
 
|pinnule arrangement=adjacent
 
|pinnule length or size=longer
 
|pinnule margin architecture or shape=serrate
 
|pinnule orientation=basiscopic
 
|scale arrangement=scattered
 
|scale coloration=tan
 
|sorus position=midway
 
|tooth architecture or shape=spiny
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Dryopteris]]
 
-->[[Category:Treatment]][[Category:Dryopteris]]

Latest revision as of 22:18, 20 February 2024

Leaves monomorphic, dying in winter, 15–75 × 10–30 cm. Petiole 1/4–1/3 length of leaf, scaly at least at base; scales scattered, tan. Blade light green, ovate-lanceolate, 2–3-pinnate-pinnatifid, herbaceous, not glandular. Pinnae ± in plane of blade, lance-oblong; basal pinnae lanceolate-deltate, slightly reduced, basal pinnules usually longer than adjacent pinnules, basal basiscopic pinnule longer than basal acroscopic pinnule; pinnule margins serrate, teeth spiny. Sori midway between midvein and margin of segments. Indusia lacking glands. 2n = 164.


Habitat: Swampy woods, moist wooded slopes, stream banks, and conifer plantations
Elevation: 0–1200 m

Distribution

V2 507-distribution-map.gif

Alta., B.C., Man., N.B., Nfld. and Labr., N.W.T., N.S., Ont., P.E.I., Que., Sask., Yukon, Ark., Conn., Del., Idaho, Ill., Ind., Iowa, Ky., Maine, Md., Mass., Mich., Minn., Mo., Mont., Nebr., N.H., N.J., N.Y., N.C., N.Dak., Ohio, Pa., R.I., S.C., Tenn., Vt., Va., Wash., W.Va., Wis., Eurasia.

Discussion

Dryopteris carthusiana is tetraploid. Dryopteris intermedia is one parent, as indicated by chromosome pairing in their hybrid D. × triploidea Wherry. The other parent is the hypothetical missing ancestral species " D. semicristata " (see discussion for D. cristata). Dryopteris carthusiana hybridizes with five species; hybrids can be separated from D. intermedia by the lack of glandular hairs and by having 2-pinnate leaves.

Selected References

None.

Lower Taxa

None.
... more about "Dryopteris carthusiana"
James D. Montgomery +  and Warren H. Wagner Jr. +
(Villars) H. P. Fuchs +
Polypodium carthusianum +
Spinulose wood fern +, toothed wood fern +  and dryoptère de cartheuser +
Alta. +, B.C. +, Man. +, N.B. +, Nfld. +, N.W.T. +, N.S. +, Ont. +, P.E.I. +, Que. +, Sask. +, Yukon +, Ark. +, Conn. +, Del. +, Idaho +, Ill. +, Ind. +, Iowa +, Ky. +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, Mo. +, Mont. +, Nebr. +, N.H. +, N.J. +, N.Y. +, N.C. +, N.Dak. +, Ohio +, Pa. +, R.I. +, S.C. +, Tenn. +, Vt. +, Va. +, Wash. +, W.Va. +, Wis. +  and Eurasia. +
0–1200 m +
Swampy woods, moist wooded slopes, stream banks, and conifer plantations +
Bull. Soc. Bot. France +
Dryopteris austriaca var. spinulosa +, Dryopteris spinulosa +  and Polypodium spinulosum +
Dryopteris carthusiana +
Dryopteris +
species +