Difference between revisions of "Woodsia alpina"

(Bolton) Gray

Nat. Arr. Brit. Pl. 2: 17. 1822.

Common names: Alpine cliff fern woodsie alpine
Basionym: Acrostichum alpinum Bolton Fil. Brit. 2: 76, plate 42. 1790
Synonyms: Woodsia alpina var. bellii Lawson Woodsia bellii (Liljeblad) R. Brown Woodsia hyperborea (Linnaeus) R. Brown Woodsia ilvensis var. alpina (Bolton) Watt
Treatment appears in FNA Volume 2.
FNA>Volume Importer
 
m (Fixed Nfld. and Labr. distribution to match printed version.)
 
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|publications={{Treatment/Publication
 
|publications={{Treatment/Publication
 
|title=Nat. Arr. Brit. Pl.
 
|title=Nat. Arr. Brit. Pl.
|place=2: 17. 1821
+
|place=2: 17. 1822
|year=1821
+
|year=1822
 
}}
 
}}
 
|common_names=Alpine cliff fern;woodsie alpine
 
|common_names=Alpine cliff fern;woodsie alpine
|basionyms={{Treatment/ID/Synonym
+
|basionyms={{Treatment/ID/Basionym
 
|name=Acrostichum alpinum
 
|name=Acrostichum alpinum
 
|authority=Bolton
 
|authority=Bolton
 +
|rank=species
 +
|publication_title=Fil. Brit.
 +
|publication_place=2: 76, plate 42. 1790
 
}}
 
}}
 
|synonyms={{Treatment/ID/Synonym
 
|synonyms={{Treatment/ID/Synonym
 
|name=Woodsia alpina var. bellii
 
|name=Woodsia alpina var. bellii
 
|authority=Lawson
 
|authority=Lawson
}}{{Treatment/ID/Synonym
+
|rank=variety
 +
}} {{Treatment/ID/Synonym
 
|name=Woodsia bellii
 
|name=Woodsia bellii
 
|authority=(Liljeblad) R. Brown
 
|authority=(Liljeblad) R. Brown
}}{{Treatment/ID/Synonym
+
|rank=species
 +
}} {{Treatment/ID/Synonym
 
|name=Woodsia hyperborea
 
|name=Woodsia hyperborea
 
|authority=(Linnaeus) R. Brown
 
|authority=(Linnaeus) R. Brown
}}{{Treatment/ID/Synonym
+
|rank=species
 +
}} {{Treatment/ID/Synonym
 
|name=Woodsia ilvensis var. alpina
 
|name=Woodsia ilvensis var. alpina
 
|authority=(Bolton) Watt
 
|authority=(Bolton) Watt
 +
|rank=variety
 
}}
 
}}
 
|hierarchy=Dryopteridaceae;Woodsia;Woodsia alpina
 
|hierarchy=Dryopteridaceae;Woodsia;Woodsia alpina
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}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="stem architecture or arrangement;stem orientation;stem orientation;stem orientation;petiole base duration"><b>Stems </b>compact, erect to ascending, with cluster of persistent petiole bases of ± equal length;</span> <span class="statement" id="st-d0_s1" data-properties="scale coloration;scale shape">scales uniformly brown, lanceolate.</span> <span class="statement" id="st-d0_s2" data-properties="leaf length;leaf width"><b>Leaves </b>2.5–20 × 0.5–2.5 cm.</span> <span class="statement" id="st-d0_s3" data-properties="petiole coloration;petiole coloration;petiole architecture;petiole fragility;node shape"><b>Petiole </b>reddish-brown or dark purple when mature, articulate above base at swollen node, relatively brittle and easily shattered.</span> <span class="statement" id="st-d0_s4" data-properties="blade shape;blade shape;blade shape;blade shape;gland count;gland coating"><b>Blade </b>linear to narrowly lanceolate, usually pinnate-pinnatifid proximally, lacking glands, never viscid;</span> <span class="statement" id="st-d0_s5" data-properties="hair arrangement">rachis with widely scattered hairs and scales.</span> <span class="statement" id="st-d0_s6" data-properties="pinna shape;pinna shape;pinna shape;pinna length or size;pinna shape;apex shape;apex shape"><b>Pinnae </b>ovatelanceolate to deltate, longer than wide, abruptly tapered to a rounded or broadly acute apex;</span> <span class="statement" id="st-d0_s7" data-properties="pair count">largest pinnae with 1–3 pairs of pinnules;</span> <span class="statement" id="st-d0_s8" data-properties="hair arrangement;scale arrangement or course or shape;adaxial surface pubescence">abaxial surface with isolated hairs and linear scales, adaxial surface glabrous.</span> <span class="statement" id="st-d0_s9" data-properties="pinnule shape;pinnule shape"><b>Pinnules </b>entire or broadly crenate;</span> <span class="statement" id="st-d0_s10" data-properties="margin reflectance;margin width;cilium count;cilium arrangement;projection count;projection coloration or reflectance">margins nonlustrous, thin, with occasional isolated cilia, lacking translucent projections.</span> <span class="statement" id="st-d0_s11" data-properties="vein tip size;hydathode coloration;hydathode prominence"><b>Vein </b>tips often enlarged to form whitish hydathodes visible adaxially.</span> <span class="statement" id="st-d0_s12" data-properties="indusium architecture or arrangement;indusium count;indusium length or size;segment size or width;segment shape;sporangium life cycle"><b>Indusia </b>of narrow, hairlike segments, these uniseriate throughout, composed of cells many times longer than wide, usually surpassing mature sporangia.</span> <span class="statement" id="st-d0_s13" data-properties=""><b>Spores </b>averaging 46–53 µm.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact, erect to ascending, with cluster of persistent petiole bases of ± equal length; scales uniformly brown, lanceolate. <b>Leaves</b> 2.5–20 × 0.5–2.5 cm. <b>Petiole</b> reddish brown or dark purple when mature, articulate above base at swollen node, relatively brittle and easily shattered. <b>Blade</b> linear to narrowly lanceolate, usually pinnate-pinnatifid proximally, lacking glands, never viscid; rachis with widely scattered hairs and scales. <b>Pinnae</b> ovate-lanceolate to deltate, longer than wide, abruptly tapered to a rounded or broadly acute apex; largest pinnae with 1–3 pairs of pinnules; abaxial surface with isolated hairs and linear scales, adaxial surface glabrous. <b>Pinnules</b> entire or broadly crenate; margins nonlustrous, thin, with occasional isolated cilia, lacking translucent projections. <b>Vein</b> tips often enlarged to form whitish hydathodes visible adaxially. <b>Indusia</b> of narrow, hairlike segments, these uniseriate throughout, composed of cells many times longer than wide, usually surpassing mature sporangia. <b>Spores</b> averaging 46–53 µm.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
 +
|phenology=Sporulating summer–early fall.
 
|habitat=Crevices and ledges on cliffs (occasionally on rocky slopes), mostly slaty and calcareous rocks
 
|habitat=Crevices and ledges on cliffs (occasionally on rocky slopes), mostly slaty and calcareous rocks
 
|elevation=0–1500 m
 
|elevation=0–1500 m
|distribution=Greenland;B.C.;Man.;N.B.;Nfld.;N.W.T.;N.S.;Ont.;Que.;Sask.;Yukon;Alaska;Maine;Mich.;Minn.;N.H.;N.Y.;Vt.;n Eurasia.
+
|distribution=Greenland;B.C.;Man.;N.B.;Nfld. and Labr.;N.W.T.;N.S.;Nunavut;Ont.;Que.;Sask.;Yukon;Alaska;Maine;Mich.;Minn.;N.H.;N.Y.;Vt.;n Eurasia.
|discussion=<p>Isozyme studies confirm the longstanding hypothesis that Woodsia alpina is an allotetraploid derived from hybridization between W. glabella and W. ilvensis (see reticulogram). Considerable disagreement exists concerning the chromosome number of W. alpina, but 2n = 160 seems most likely, given the numbers reported for the two parental species. Hybrids between W. alpina and W. ilvensis have been reported from both Europe and North America. These morphologically intermediate triploids with malformed spores have been called W. × gracilis (Lawson) Butters.</p>
+
|discussion=<p>Isozyme studies confirm the longstanding hypothesis that <i>Woodsia alpina</i> is an allotetraploid derived from hybridization between <i>W. glabella</i> and <i>W. ilvensis</i> (see reticulogram). Considerable disagreement exists concerning the chromosome number of <i>W. alpina</i>, but 2n = 160 seems most likely, given the numbers reported for the two parental species. Hybrids between <i>W. alpina</i> and <i>W. ilvensis</i> have been reported from both Europe and North America. These morphologically intermediate triploids with malformed spores have been called W. × gracilis (Lawson) Butters.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Woodsia alpina
 
name=Woodsia alpina
|author=
 
 
|authority=(Bolton) Gray
 
|authority=(Bolton) Gray
 
|rank=species
 
|rank=species
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|basionyms=Acrostichum alpinum
 
|basionyms=Acrostichum alpinum
 
|family=Dryopteridaceae
 
|family=Dryopteridaceae
 +
|phenology=Sporulating summer–early fall.
 
|habitat=Crevices and ledges on cliffs (occasionally on rocky slopes), mostly slaty and calcareous rocks
 
|habitat=Crevices and ledges on cliffs (occasionally on rocky slopes), mostly slaty and calcareous rocks
 
|elevation=0–1500 m
 
|elevation=0–1500 m
|distribution=Greenland;B.C.;Man.;N.B.;Nfld.;N.W.T.;N.S.;Ont.;Que.;Sask.;Yukon;Alaska;Maine;Mich.;Minn.;N.H.;N.Y.;Vt.;n Eurasia.
+
|distribution=Greenland;B.C.;Man.;N.B.;Nfld. and Labr.;N.W.T.;N.S.;Nunavut;Ont.;Que.;Sask.;Yukon;Alaska;Maine;Mich.;Minn.;N.H.;N.Y.;Vt.;n Eurasia.
 
|reference=None
 
|reference=None
 
|publication title=Nat. Arr. Brit. Pl.
 
|publication title=Nat. Arr. Brit. Pl.
|publication year=1821
+
|publication year=1822
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_48.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_48.xml
 
|genus=Woodsia
 
|genus=Woodsia
 
|species=Woodsia alpina
 
|species=Woodsia alpina
|adaxial surface pubescence=glabrous
 
|apex shape=acute;rounded
 
|blade shape=pinnate-pinnatifid;linear;narrowly lanceolate
 
|cilium arrangement=isolated
 
|cilium count=occasional
 
|gland coating=viscid
 
|gland count=lacking
 
|hair arrangement=isolated;scattered
 
|hydathode coloration=whitish
 
|hydathode prominence=visible
 
|indusium architecture or arrangement=uniseriate
 
|indusium count=many
 
|indusium length or size=longer than wide
 
|leaf length=2.5cm;20cm
 
|leaf width=0.5cm;2.5cm
 
|margin reflectance=nonlustrous
 
|margin width=thin
 
|node shape=swollen
 
|pair count=1;3
 
|petiole architecture=articulate
 
|petiole base duration=persistent
 
|petiole coloration=dark purple;reddish-brown
 
|petiole fragility=brittle
 
|pinna length or size=longer than wide
 
|pinna shape=tapered;ovatelanceolate;deltate
 
|pinnule shape=crenate;entire
 
|projection coloration or reflectance=translucent
 
|projection count=lacking
 
|scale arrangement or course or shape=linear
 
|scale coloration=brown
 
|scale shape=lanceolate
 
|segment shape=hairlike
 
|segment size or width=narrow
 
|sporangium life cycle=mature
 
|stem architecture or arrangement=compact
 
|stem orientation=erect;ascending
 
|vein tip size=enlarged
 
 
}}<!--
 
}}<!--
  
-->[[Category:Treatment]][[Category:Woodsia]]
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-->
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 +
[[Category:Treatment]]
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[[Category:Woodsia]]
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[[Category:Revised Since Print]]

Latest revision as of 21:39, 20 February 2024

Stems compact, erect to ascending, with cluster of persistent petiole bases of ± equal length; scales uniformly brown, lanceolate. Leaves 2.5–20 × 0.5–2.5 cm. Petiole reddish brown or dark purple when mature, articulate above base at swollen node, relatively brittle and easily shattered. Blade linear to narrowly lanceolate, usually pinnate-pinnatifid proximally, lacking glands, never viscid; rachis with widely scattered hairs and scales. Pinnae ovate-lanceolate to deltate, longer than wide, abruptly tapered to a rounded or broadly acute apex; largest pinnae with 1–3 pairs of pinnules; abaxial surface with isolated hairs and linear scales, adaxial surface glabrous. Pinnules entire or broadly crenate; margins nonlustrous, thin, with occasional isolated cilia, lacking translucent projections. Vein tips often enlarged to form whitish hydathodes visible adaxially. Indusia of narrow, hairlike segments, these uniseriate throughout, composed of cells many times longer than wide, usually surpassing mature sporangia. Spores averaging 46–53 µm.


Phenology: Sporulating summer–early fall.
Habitat: Crevices and ledges on cliffs (occasionally on rocky slopes), mostly slaty and calcareous rocks
Elevation: 0–1500 m

Distribution

V2 48-distribution-map.gif

Greenland, B.C., Man., N.B., Nfld. and Labr., N.W.T., N.S., Nunavut, Ont., Que., Sask., Yukon, Alaska, Maine, Mich., Minn., N.H., N.Y., Vt., n Eurasia.

Discussion

Isozyme studies confirm the longstanding hypothesis that Woodsia alpina is an allotetraploid derived from hybridization between W. glabella and W. ilvensis (see reticulogram). Considerable disagreement exists concerning the chromosome number of W. alpina, but 2n = 160 seems most likely, given the numbers reported for the two parental species. Hybrids between W. alpina and W. ilvensis have been reported from both Europe and North America. These morphologically intermediate triploids with malformed spores have been called W. × gracilis (Lawson) Butters.

Selected References

None.

Lower Taxa

None.
... more about "Woodsia alpina"
Michael D. Windham +
(Bolton) Gray +
Acrostichum alpinum +
Alpine cliff fern +  and woodsie alpine +
Greenland +, B.C. +, Man. +, N.B. +, Nfld. and Labr. +, N.W.T. +, N.S. +, Nunavut +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Maine +, Mich. +, Minn. +, N.H. +, N.Y. +, Vt. +  and n Eurasia. +
0–1500 m +
Crevices and ledges on cliffs (occasionally on rocky slopes), mostly slaty and calcareous rocks +
Sporulating summer–early fall. +
Nat. Arr. Brit. Pl. +
Woodsia alpina var. bellii +, Woodsia bellii +, Woodsia hyperborea +  and Woodsia ilvensis var. alpina +
Woodsia alpina +
species +