Difference between revisions of "Asplenium bradleyi"

D. C. Eaton

Bull. Torrey Bot. Club 4: 11. 1873.

Common names: Bradley's spleenwort
Endemic
Synonyms: Asplenium stotleri Wherry
Treatment appears in FNA Volume 2.
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|common_names=Bradley's spleenwort
 
|common_names=Bradley's spleenwort
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|special_status={{Treatment/ID/Special_status
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|code=E
 +
|label=Endemic
 +
}}
 
|basionyms=
 
|basionyms=
 
|synonyms={{Treatment/ID/Synonym
 
|synonyms={{Treatment/ID/Synonym
 
|name=Asplenium stotleri
 
|name=Asplenium stotleri
 
|authority=Wherry
 
|authority=Wherry
 +
|rank=species
 
}}
 
}}
 
|hierarchy=Aspleniaceae;Asplenium;Asplenium bradleyi
 
|hierarchy=Aspleniaceae;Asplenium;Asplenium bradleyi
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}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="root reproduction"><b>Roots </b>not proliferous.</span> <span class="statement" id="st-d0_s1" data-properties="stem orientation;stem orientation;stem orientation;stem architecture"><b>Stems </b>short-creeping to ascending, occasionally branched;</span> <span class="statement" id="st-d0_s2" data-properties="scale coloration;scale coloration;scale coloration;scale shape;scale atypical length;scale length;scale width;margin architecture or shape;margin architecture or shape">scales dark reddish to brown throughout, narrowly deltate, (2–) 3–5 × 0.2–0.4 mm, margins entire or shallowly dentate.</span> <span class="statement" id="st-d0_s3" data-properties="leaf architecture"><b>Leaves </b>monomorphic.</span> <span class="statement" id="st-d0_s4" data-properties="petiole coloration;petiole coloration;petiole reflectance;petiole atypical some measurement;petiole some measurement;petiole length"><b>Petiole </b>reddish or purplish brown throughout, lustrous, 1–10 (–13) cm, 1/3–3/4 length of blade;</span> <span class="statement" id="st-d0_s5" data-properties="scale coloration;scale shape">indument of brown, narrowly lanceolate scales at base, grading into hairs.</span> <span class="statement" id="st-d0_s6" data-properties="blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade atypical length;blade length;blade width;blade width;blade width;blade width;blade pubescence"><b>Blade </b>narrowly oblong to lanceolate, pinnate-pinnatifid to 2-pinnate, 2–17 (–20) × 1–6 cm, thin to moderately thick, sparsely pubescent;</span> <span class="statement" id="st-d0_s7" data-properties="base shape;base shape">base truncate or obtuse;</span> <span class="statement" id="st-d0_s8" data-properties="apex shape;apex architecture">apex acute, not rooting.</span> <span class="statement" id="st-d0_s9" data-properties="rachis coloration;rachis coloration;rachis coloration;rachis coloration;rachis coloration;rachis coloration;rachis reflectance;rachis pubescence"><b>Rachis </b>reddish or purplish brown proximally, fading to green in distal 1/3–2/3, lustrous, sparsely pubescent.</span> <span class="statement" id="st-d0_s10" data-properties="pinna shape;pinna shape;pinna shape;pinna shape;pinna shape;pair atypical count;pair atypical count;pair count"><b>Pinnae </b>in (3–) 5–15 (–25) pairs, ovate, obovate to lanceolate or deltate-lanceolate;</span> <span class="statement" id="st-d0_s11" data-properties="medial pinna length;medial pinna width">medial pinnae 6–40 × 3–10 mm;</span> <span class="statement" id="st-d0_s12" data-properties="base shape;base shape;base shape">base truncate to obliquely obtuse;</span> <span class="statement" id="st-d0_s13" data-properties="margin shape;margin shape;margin shape">margins dentate to denticulate;</span> <span class="statement" id="st-d0_s14" data-properties="apex shape;apex shape">apex acute or rounded.</span> <span class="statement" id="st-d0_s15" data-properties="vein fusion;vein prominence"><b>Veins </b>free, barely evident.</span> <span class="statement" id="st-d0_s16" data-properties="sorus count;pair count;side orientation;side orientation"><b>Sori </b>3 to numerous pairs per pinna, on both basiscopic and acroscopic sides.</span> <span class="statement" id="st-d0_s17" data-properties=""><b>Spores </b>64 per sporangium.</span> <span class="statement" id="st-d0_s18" data-properties="spore count;2n chromosome count">2n = 144.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Roots </b>not proliferous. <b>Stems</b> short-creeping to ascending, occasionally branched; scales dark reddish to brown throughout, narrowly deltate, (2–)3–5 × 0.2–0.4 mm, margins entire or shallowly dentate. <b>Leaves</b> monomorphic. <b>Petiole</b> reddish or purplish brown throughout, lustrous, 1–10(–13) cm, 1/3–3/4 length of blade; indument of brown, narrowly lanceolate scales at base, grading into hairs. <b>Blade</b> narrowly oblong to lanceolate, pinnate-pinnatifid to 2-pinnate, 2–17(–20) × 1–6 cm, thin to moderately thick, sparsely pubescent; base truncate or obtuse; apex acute, not rooting. <b>Rachis</b> reddish or purplish brown proximally, fading to green in distal 1/3–2/3, lustrous, sparsely pubescent. <b>Pinnae</b> in (3–)5–15(–25) pairs, ovate, obovate to lanceolate or deltate-lanceolate; medial pinnae 6–40 × 3–10 mm; base truncate to obliquely obtuse; margins dentate to denticulate; apex acute or rounded. <b>Veins</b> free, barely evident. <b>Sori</b> 3 to numerous pairs per pinna, on both basiscopic and acroscopic sides. <b>Spores</b> 64 per sporangium. <b>2n</b> = 144.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
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|elevation=0–1000 m
 
|elevation=0–1000 m
 
|distribution=Ala.;Ark.;Ga.;Ill.;Ky.;Md.;Mo.;N.J.;N.Y.;N.C.;Ohio;Okla.;Pa.;S.C.;Tenn.;Va.;W.Va.
 
|distribution=Ala.;Ark.;Ga.;Ill.;Ky.;Md.;Mo.;N.J.;N.Y.;N.C.;Ohio;Okla.;Pa.;S.C.;Tenn.;Va.;W.Va.
|discussion=<p>Asplenium bradleyi is a morphologically variable species, the allotetraploid derivative of A. montanum × platyneuron (W. H. Wagner Jr. 1954; D. M. Smith and D. A. Levin 1963; C. R. Werth et al. 1985). The sterile diploid form of A. bradleyi has been collected twice in nature (W. H. Wagner Jr. et al. 1973; A. M. Evans 1988), and isozyme studies indicate that the allotetraploid has had a polytopic origin (C. R. Werth et al. 1985b). Occurring rarely to locally in the Appalachian region, A. bradleyi overlaps with both progenitor taxa, but it is fairly frequent in the Ozark and Ouachita region where A. montanum is absent. Sterile hybrids with A. pinnatifidum (A. × gravesii), A. montanum (A. × wherryi D. M. Smith et al.), and A. platyneuron are known from nature.</p>
+
|discussion=<p><i>Asplenium bradleyi</i> is a morphologically variable species, the allotetraploid derivative of <i>A. montanum</i> × platyneuron (W. H. Wagner Jr. 1954; D. M. Smith and D. A. Levin 1963; C. R. Werth et al. 1985). The sterile diploid form of <i>A. bradleyi</i> has been collected twice in nature (W. H. Wagner Jr. et al. 1973; A. M. Evans 1988), and isozyme studies indicate that the allotetraploid has had a polytopic origin (C. R. Werth et al. 1985b). Occurring rarely to locally in the Appalachian region, <i>A. bradleyi</i> overlaps with both progenitor taxa, but it is fairly frequent in the Ozark and Ouachita region where <i>A. montanum</i> is absent. Sterile hybrids with <i>A. pinnatifidum</i> (A. × gravesii), <i>A. montanum</i> (A. × wherryi D. M. Smith et al.), and <i>A. platyneuron</i> are known from nature.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Asplenium bradleyi
 
name=Asplenium bradleyi
|author=
 
 
|authority=D. C. Eaton
 
|authority=D. C. Eaton
 
|rank=species
 
|rank=species
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|publication title=Bull. Torrey Bot. Club
 
|publication title=Bull. Torrey Bot. Club
 
|publication year=1873
 
|publication year=1873
|special status=
+
|special status=Endemic
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_326.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_326.xml
 
|genus=Asplenium
 
|genus=Asplenium
 
|species=Asplenium bradleyi
 
|species=Asplenium bradleyi
|2n chromosome count=144
 
|apex architecture=not rooting
 
|apex shape=rounded;acute;acute
 
|base shape=truncate;obliquely obtuse
 
|blade atypical length=17cm;20cm
 
|blade length=2cm;17cm
 
|blade pubescence=pubescent
 
|blade shape=narrowly oblong;lanceolate pinnate-pinnatifid
 
|blade width=thin;moderately thick
 
|leaf architecture=monomorphic
 
|margin architecture or shape=dentate;entire
 
|margin shape=dentate;denticulate
 
|medial pinna length=6mm;40mm
 
|medial pinna width=3mm;10mm
 
|pair atypical count=15;25
 
|pair count=numerous;5;15
 
|petiole atypical some measurement=10cm;13cm
 
|petiole coloration=purplish brown;reddish
 
|petiole length=1/3 length of blade;3/4 length of blade
 
|petiole reflectance=lustrous
 
|petiole some measurement=1cm;10cm
 
|pinna shape=obovate;lanceolate or deltate-lanceolate
 
|rachis coloration=fading;green
 
|rachis pubescence=pubescent
 
|rachis reflectance=lustrous
 
|root reproduction=not proliferous
 
|scale atypical length=2mm;3mm
 
|scale coloration=brown;dark reddish;brown
 
|scale length=3mm;5mm
 
|scale shape=lanceolate;deltate
 
|scale width=0.2mm;0.4mm
 
|side orientation=acroscopic;basiscopic
 
|sorus count=3
 
|spore count=64
 
|stem architecture=branched
 
|stem orientation=short-creeping;ascending
 
|vein fusion=free
 
|vein prominence=evident
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Asplenium]]
 
-->[[Category:Treatment]][[Category:Asplenium]]

Latest revision as of 20:22, 5 November 2020

Roots not proliferous. Stems short-creeping to ascending, occasionally branched; scales dark reddish to brown throughout, narrowly deltate, (2–)3–5 × 0.2–0.4 mm, margins entire or shallowly dentate. Leaves monomorphic. Petiole reddish or purplish brown throughout, lustrous, 1–10(–13) cm, 1/3–3/4 length of blade; indument of brown, narrowly lanceolate scales at base, grading into hairs. Blade narrowly oblong to lanceolate, pinnate-pinnatifid to 2-pinnate, 2–17(–20) × 1–6 cm, thin to moderately thick, sparsely pubescent; base truncate or obtuse; apex acute, not rooting. Rachis reddish or purplish brown proximally, fading to green in distal 1/3–2/3, lustrous, sparsely pubescent. Pinnae in (3–)5–15(–25) pairs, ovate, obovate to lanceolate or deltate-lanceolate; medial pinnae 6–40 × 3–10 mm; base truncate to obliquely obtuse; margins dentate to denticulate; apex acute or rounded. Veins free, barely evident. Sori 3 to numerous pairs per pinna, on both basiscopic and acroscopic sides. Spores 64 per sporangium. 2n = 144.


Habitat: Acidic rocks, usually on steep ledges
Elevation: 0–1000 m

Distribution

V2 326-distribution-map.gif

Ala., Ark., Ga., Ill., Ky., Md., Mo., N.J., N.Y., N.C., Ohio, Okla., Pa., S.C., Tenn., Va., W.Va.

Discussion

Asplenium bradleyi is a morphologically variable species, the allotetraploid derivative of A. montanum × platyneuron (W. H. Wagner Jr. 1954; D. M. Smith and D. A. Levin 1963; C. R. Werth et al. 1985). The sterile diploid form of A. bradleyi has been collected twice in nature (W. H. Wagner Jr. et al. 1973; A. M. Evans 1988), and isozyme studies indicate that the allotetraploid has had a polytopic origin (C. R. Werth et al. 1985b). Occurring rarely to locally in the Appalachian region, A. bradleyi overlaps with both progenitor taxa, but it is fairly frequent in the Ozark and Ouachita region where A. montanum is absent. Sterile hybrids with A. pinnatifidum (A. × gravesii), A. montanum (A. × wherryi D. M. Smith et al.), and A. platyneuron are known from nature.

Selected References

None.

Lower Taxa

None.
... more about "Asplenium bradleyi"
Warren H. Wagner Jr. +, Robbin C. Moran +  and Charles R. Werth +
D. C. Eaton +
Bradley's spleenwort +
Ala. +, Ark. +, Ga. +, Ill. +, Ky. +, Md. +, Mo. +, N.J. +, N.Y. +, N.C. +, Ohio +, Okla. +, Pa. +, S.C. +, Tenn. +, Va. +  and W.Va. +
0–1000 m +
Acidic rocks, usually on steep ledges +
Bull. Torrey Bot. Club +
Asplenium stotleri +
Asplenium bradleyi +
Asplenium +
species +