Difference between revisions of "Echinocactus"

Link & Otto

Verh. Vereins Beförd. Gartenbaues Königl. Preuss. Staaten 3: 420. 1827.

Common names: Barrel cactus eagle-claw cactus echinocactus
Etymology: Greek echinos, hedgehog, an d Cactus, an old genus name
Synonyms: Homalocephala Britton & Rose
Treatment appears in FNA Volume 4. Treatment on page 188. Mentioned on page 97, 207.
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|name=Homalocephala
 
|name=Homalocephala
 
|authority=Britton & Rose
 
|authority=Britton & Rose
 +
|rank=genus
 
}}
 
}}
 
|hierarchy=Cactaceae;Cactaceae subfam. Cactoideae;Echinocactus
 
|hierarchy=Cactaceae;Cactaceae subfam. Cactoideae;Echinocactus
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--><span class="statement" id="st-d0_s0" data-properties="plant orientation;plant architecture;plant architecture;plant location;mound architecture or arrangement;branch count"><b>Plants </b>erect, branched or unbranched, forming compact mounds of 30+ branches, not deep-seated in substrate (or deep-seated in E. texensis and, if plants immature or at high altitude, in E. horizonthalonius).</span> <span class="statement" id="st-d0_s1" data-properties="taproot density;taproot height or length or size"><b>Roots </b>diffuse or short taproots.</span> <span class="statement" id="st-d0_s2" data-properties="stem architecture;stem coloration;stem coloration;stem coloration;stem coloration;stem coloration;stem coloration;stem shape;stem shape;stem shape;stem shape;stem shape;stem atypical length;stem length;stem atypical width;stem width;apical region pubescence"><b>Stems </b>unsegmented, gray-blue, gray-green, yellow-green, or grass green, flat-topped spheric to short cylindric, 4–40 (–45) [–250] × 8–30 [–80] cm, apical region appearing copiously woolly (shortly velvety in E. texensis) [glabrous];</span> <span class="statement" id="st-d0_s3" data-properties="rib atypical count;rib atypical count;rib count;rib prominence;rib course;rib course;rib shape;rib shape;rib shape;rib crest architecture;rib crest size">ribs (7–) 8–27 [–60+], very prominent, straight (or only slightly undulate), sometimes helically curving around stems, broadly rounded to nearly keeled, rib crests uninterrupted or ± constricted between areoles;</span> <span class="statement" id="st-d0_s4" data-properties="areole arrangement;areole arrangement;areole shape;areole shape;areole shape;portion reproduction;adaxial portion height or length or size;adaxial portion width;adaxial portion arrangement;spine arrangement">areoles widely spaced or confluent with age, nearly circular to oblong, with fertile portion as short, broad adaxial prolongation confluent with spine cluster;</span> <span class="statement" id="st-d0_s5" data-properties="areolar gland count">areolar glands absent;</span> <span class="statement" id="st-d0_s6" data-properties="cortex texture;cortex coating;pith texture;pith coating">cortex and pith hard, not mucilaginous.</span> <span class="statement" id="st-d0_s7" data-properties="spine atypical count;spine count;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine shape;spine texture;spine fragility"><b>Spines </b>(5–) 7–19 per areole, straw colored, pink, red, gray, tan, or brown, conspicuously annulate-ridged, rigid, stiff;</span> <span class="statement" id="st-d0_s8" data-properties="spine arrangement;spine count;spine course;spine course;spine course;spine some measurement">radial spines 5–14 per areole, straight to curved, 2–5 cm;</span> <span class="statement" id="st-d0_s9" data-properties="central spine count;central spine course;central spine course;central spine course;central spine shape;central spine shape;central spine shape;central spine shape;central spine shape">central spines 1–4 per areole, straight to curved, terete, flattened, or abaxially ridged.</span> <span class="statement" id="st-d0_s10" data-properties="flower duration;flower shape;flower shape;flower shape;adaxial extension height or length or size"><b>Flowers </b>diurnal, near stem apex, near adaxial edges of short adaxial extensions of areoles, broadly funnelform to nearly salverform (more narrowly funnelform in E. texensis);</span> <span class="statement" id="st-d0_s11" data-properties="outer tepals margin architecture or shape;outer tepals margin architecture or shape">outer tepals margins entire, apically spinose;</span> <span class="statement" id="st-d0_s12" data-properties="inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal length;inner tepal width;margin architecture or shape;margin architecture or shape;margin shape;margin architecture or relief">inner tepals yellow or pink to magenta, 2.4–3.2 × 0.3–1.5 cm, margins entire, serrate, toothed, or erose;</span> <span class="statement" id="st-d0_s13" data-properties="ovary architecture or pubescence;ovary architecture;ovary pubescence;hair coloration;hair coloration">ovary scaly, spineless, copiously woolly, with white or pale tan hairs from areoles hiding ovary and flower tube at anthesis;</span> <span class="statement" id="st-d0_s14" data-properties="scale count;margin shape;margin shape;margin architecture or shape">scales 8–60, margins entire or fimbriate, spine-tipped;</span> <span class="statement" id="st-d0_s15" data-properties="stigma lobe atypical count;stigma lobe count;stigma lobe coloration;stigma lobe coloration;stigma lobe coloration;stigma lobe coloration;stigma lobe coloration;stigma lobe some measurement">stigma lobes 6–14 (–17), yellow, pink, or olive, 1–4.5 mm.</span> <span class="statement" id="st-d0_s16" data-properties="fruit coloration;fruit coloration;fruit coloration;fruit shape;fruit shape;fruit shape;fruit shape;fruit length;fruit atypical width;fruit width;fruit condition or texture;fruit count;pore position;pore life cycle;pore dehiscence;pore dehiscence;pore dehiscence;pore dehiscence;pore position;pore life cycle"><b>Fruits </b>either indehiscent or rupturing irregularly, or tardily dehiscent through basal abscission pore, whitish tan to pinkish (bright red in E. texensis), spheric to ovoid or ovoid-cylindric, 10–50 × 10–40 [–100] mm, usually nearly dry (strongly succulent in E. texensis), many scaled;</span> <span class="statement" id="st-d0_s17" data-properties="axil pubescence;axil architecture;distal scale architecture or shape">axils of scales copiously woolly (wool hiding surface of fruits except in E. texensis), spineless, distal scales spine-tipped;</span> <span class="statement" id="st-d0_s18" data-properties="pulp coloration">pulp red;</span> <span class="statement" id="st-d0_s19" data-properties="floral remnant duration">floral remnant persistent.</span> <span class="statement" id="st-d0_s20" data-properties="seed coloration;seed coloration;seed coloration;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed some measurement;seed reflectance;seed reflectance"><b>Seeds </b>reddish-brown to black, spheric, or subreniform to obovoid, 2.4–4.7 mm, shiny or dull;</span> <span class="statement" id="st-d0_s21" data-properties="">testa cells convex or flat.</span> <span class="statement" id="st-d0_s22" data-properties="testa cell shape;testa cell shape;x chromosome count">x = 11.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>erect, branched or unbranched, forming compact mounds of 30+ branches, not deep-seated in substrate (or deep-seated in <i>E. texensis</i> and, if plants immature or at high altitude, in <i>E. horizonthalonius</i>). <b>Roots</b> diffuse or short taproots. <b>Stems</b> unsegmented, gray-blue, gray-green, yellow-green, or grass green, flat-topped spheric to short cylindric, 4–40(–45)[–250] × 8–30[–80] cm, apical region appearing copiously woolly (shortly velvety in <i>E. texensis</i>) [glabrous]; ribs (7–)8–27[–60+], very prominent, straight (or only slightly undulate), sometimes helically curving around stems, broadly rounded to nearly keeled, rib crests uninterrupted or ± constricted between areoles; areoles widely spaced or confluent with age, nearly circular to oblong, with fertile portion as short, broad adaxial prolongation confluent with spine cluster; areolar glands absent; cortex and pith hard, not mucilaginous. <b>Spines</b> (5–)7–19 per areole, straw colored, pink, red, gray, tan, or brown, conspicuously annulate-ridged, rigid, stiff; radial spines 5–14 per areole, straight to curved, 2–5 cm; central spines 1–4 per areole, straight to curved, terete, flattened, or abaxially ridged. <b>Flowers</b> diurnal, near stem apex, near adaxial edges of short adaxial extensions of areoles, broadly funnelform to nearly salverform (more narrowly funnelform in <i>E. texensis</i>); outer tepals margins entire, apically spinose; inner tepals yellow or pink to magenta, 2.4–3.2 × 0.3–1.5 cm, margins entire, serrate, toothed, or erose; ovary scaly, spineless, copiously woolly, with white or pale tan hairs from areoles hiding ovary and flower tube at anthesis; scales 8–60, margins entire or fimbriate, spine-tipped; stigma lobes 6–14(–17), yellow, pink, or olive, 1–4.5 mm. <b>Fruits</b> either indehiscent or rupturing irregularly, or tardily dehiscent through basal abscission pore, whitish tan to pinkish (bright red in <i>E. texensis</i>), spheric to ovoid or ovoid-cylindric, 10–50 × 10–40[–100] mm, usually nearly dry (strongly succulent in <i>E. texensis</i>), many scaled; axils of scales copiously woolly (wool hiding surface of fruits except in <i>E. texensis</i>), spineless, distal scales spine-tipped; pulp red; floral remnant persistent. <b>Seeds</b> reddish brown to black, spheric, or subreniform to obovoid, 2.4–4.7 mm, shiny or dull; testa cells convex or flat. <b>x</b> = 11.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
 
|distribution=sw United States;Mexico.
 
|distribution=sw United States;Mexico.
 
|discussion=<p>Species 6 (3 in the flora).</p><!--
 
|discussion=<p>Species 6 (3 in the flora).</p><!--
--><p>With the exception of the much-branched Echinocactus polycephalus, species of Echinocactus in the flora area are almost never branched, and will die instead of producing a lateral bud when the apex is injured. Smaller than normal fruits of Echinocactus are usually aborted and hollow as the result of infestation by insect larvae (M. Chamberland, pers. comm.).</p>
+
--><p>With the exception of the much-branched <i>Echinocactus polycephalus</i>, species of <i>Echinocactus</i> in the flora area are almost never branched, and will die instead of producing a lateral bud when the apex is injured. Smaller than normal fruits of <i>Echinocactus</i> are usually aborted and hollow as the result of infestation by insect larvae (M. Chamberland, pers. comm.).</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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|publication year=1827
 
|publication year=1827
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_353.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V4/V4_353.xml
 
|subfamily=Cactaceae subfam. Cactoideae
 
|subfamily=Cactaceae subfam. Cactoideae
 
|genus=Echinocactus
 
|genus=Echinocactus
|adaxial extension height or length or size=short
 
|adaxial portion arrangement=confluent
 
|adaxial portion height or length or size=short
 
|adaxial portion width=broad
 
|apical region pubescence=woolly
 
|areolar gland count=absent
 
|areole arrangement=confluent;spaced
 
|areole shape=nearly circular;oblong
 
|axil architecture=spineless
 
|axil pubescence=woolly
 
|central spine count=1;4
 
|central spine course=straight;curved
 
|central spine shape=ridged;flattened;ridged;flattened;terete
 
|cortex coating=not mucilaginous
 
|cortex texture=hard
 
|distal scale architecture or shape=spine-tipped
 
|floral remnant duration=persistent
 
|flower duration=diurnal
 
|flower shape=broadly funnelform;nearly salverform
 
|fruit atypical width=40mm;100mm
 
|fruit coloration=whitish tan;pinkish
 
|fruit condition or texture=dry
 
|fruit count=many
 
|fruit length=10mm;50mm
 
|fruit shape=spheric;ovoid or ovoid-cylindric
 
|fruit width=10mm;40mm
 
|hair coloration=pale tan;white
 
|inner tepal coloration=pink;magenta
 
|inner tepal length=2.4cm;3.2cm
 
|inner tepal width=0.3cm;1.5cm
 
|margin architecture or relief=erose
 
|margin architecture or shape=spine-tipped;serrate;entire
 
|margin shape=fimbriate;entire;toothed
 
|mound architecture or arrangement=compact
 
|outer tepals margin architecture or shape=spinose;entire
 
|ovary architecture=spineless
 
|ovary architecture or pubescence=scaly
 
|ovary pubescence=woolly
 
|pith coating=not mucilaginous
 
|pith texture=hard
 
|plant architecture=unbranched;branched
 
|plant location=not deep-seated
 
|plant orientation=erect
 
|pore dehiscence=dehiscent;,;rupturing;indehiscent
 
|pore life cycle=abscission;abscission
 
|pore position=basal;basal
 
|portion reproduction=fertile
 
|pulp coloration=red
 
|rib atypical count=27;60
 
|rib count=8;27
 
|rib course=curving;straight
 
|rib crest architecture=uninterrupted
 
|rib crest size=constricted
 
|rib prominence=prominent
 
|rib shape=broadly rounded;nearly keeled
 
|scale count=8;60
 
|seed coloration=reddish-brown;black
 
|seed reflectance=dull;shiny
 
|seed shape=subreniform;obovoid
 
|seed some measurement=2.4mm;4.7mm
 
|spine arrangement=radial;cluster
 
|spine atypical count=5;7
 
|spine coloration=brown;tan;brown;tan;gray;red;pink;straw colored
 
|spine count=5;14
 
|spine course=straight;curved
 
|spine fragility=stiff
 
|spine shape=annulate-ridged
 
|spine some measurement=2cm;5cm
 
|spine texture=rigid
 
|stem architecture=unsegmented
 
|stem atypical length=40cm;45cm
 
|stem atypical width=30cm;80cm
 
|stem coloration=grass green;yellow-green;grass green;yellow-green;gray-green;gray-blue
 
|stem length=4cm;40cm
 
|stem shape=spheric;short cylindric
 
|stem width=8cm;30cm
 
|stigma lobe atypical count=14;17
 
|stigma lobe coloration=olive;pink;olive;pink;yellow
 
|stigma lobe count=6;14
 
|stigma lobe some measurement=1mm;4.5mm
 
|taproot density=diffuse
 
|taproot height or length or size=short
 
|testa cell shape=flat;convex
 
|x chromosome count=11
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Cactaceae subfam. Cactoideae]]
 
-->[[Category:Treatment]][[Category:Cactaceae subfam. Cactoideae]]

Latest revision as of 21:58, 5 November 2020

Plants erect, branched or unbranched, forming compact mounds of 30+ branches, not deep-seated in substrate (or deep-seated in E. texensis and, if plants immature or at high altitude, in E. horizonthalonius). Roots diffuse or short taproots. Stems unsegmented, gray-blue, gray-green, yellow-green, or grass green, flat-topped spheric to short cylindric, 4–40(–45)[–250] × 8–30[–80] cm, apical region appearing copiously woolly (shortly velvety in E. texensis) [glabrous]; ribs (7–)8–27[–60+], very prominent, straight (or only slightly undulate), sometimes helically curving around stems, broadly rounded to nearly keeled, rib crests uninterrupted or ± constricted between areoles; areoles widely spaced or confluent with age, nearly circular to oblong, with fertile portion as short, broad adaxial prolongation confluent with spine cluster; areolar glands absent; cortex and pith hard, not mucilaginous. Spines (5–)7–19 per areole, straw colored, pink, red, gray, tan, or brown, conspicuously annulate-ridged, rigid, stiff; radial spines 5–14 per areole, straight to curved, 2–5 cm; central spines 1–4 per areole, straight to curved, terete, flattened, or abaxially ridged. Flowers diurnal, near stem apex, near adaxial edges of short adaxial extensions of areoles, broadly funnelform to nearly salverform (more narrowly funnelform in E. texensis); outer tepals margins entire, apically spinose; inner tepals yellow or pink to magenta, 2.4–3.2 × 0.3–1.5 cm, margins entire, serrate, toothed, or erose; ovary scaly, spineless, copiously woolly, with white or pale tan hairs from areoles hiding ovary and flower tube at anthesis; scales 8–60, margins entire or fimbriate, spine-tipped; stigma lobes 6–14(–17), yellow, pink, or olive, 1–4.5 mm. Fruits either indehiscent or rupturing irregularly, or tardily dehiscent through basal abscission pore, whitish tan to pinkish (bright red in E. texensis), spheric to ovoid or ovoid-cylindric, 10–50 × 10–40[–100] mm, usually nearly dry (strongly succulent in E. texensis), many scaled; axils of scales copiously woolly (wool hiding surface of fruits except in E. texensis), spineless, distal scales spine-tipped; pulp red; floral remnant persistent. Seeds reddish brown to black, spheric, or subreniform to obovoid, 2.4–4.7 mm, shiny or dull; testa cells convex or flat. x = 11.

Distribution

sw United States, Mexico.

Discussion

Species 6 (3 in the flora).

With the exception of the much-branched Echinocactus polycephalus, species of Echinocactus in the flora area are almost never branched, and will die instead of producing a lateral bud when the apex is injured. Smaller than normal fruits of Echinocactus are usually aborted and hollow as the result of infestation by insect larvae (M. Chamberland, pers. comm.).

Selected References

None.

Key

1 Inner tepals yellow; plants many branched (except in very young adults); w Arizona, s Nevada, se California, Mexico Echinocactus polycephalus
1 Inner tepals pink, magenta, or proximally red and distally pink; plants unbranched (rarely few branched); c to s Arizona, New Mexico, Texas, Mexico > 2
2 Ribs (7-)8(-9), crests broadly rounded, never keeled; stems flat-topped or hemispheric to stoutly cylindric; fruit surfaces hidden by abundant wool at maturity; spines and bract tips glabrous (use lens) Echinocactus horizonthalonius
2 Ribs 13-27, crests narrow, ± keeled; stems flat-topped to hemispheric; fruit surfaces exposed at maturity, bright red; spines and bract tips densely puberulent with ± matted, strap-shaped, unicellular hairs (use at least 8× lens) Echinocactus texensis
... more about "Echinocactus"
Allan D. Zimmerman +  and Bruce D. Parfitt +
Link & Otto +
Barrel cactus +, eagle-claw cactus +  and echinocactus +
sw United States +  and Mexico. +
Greek echinos, hedgehog, an d Cactus, an old genus name +
Verh. Vereins Beförd. Gartenbaues Königl. Preuss. Staaten +
Homalocephala +
Echinocactus +
Cactaceae subfam. Cactoideae +