Difference between revisions of "Cylindropuntia spinosior"

(Engelmann) F. M. Knuth

in C. Backeberg and F. M. Knuth, Kaktus-ABC, 126. 1935.

Common names: Cane cholla walkingstick cactus
Basionym: Opuntia whipplei var. spinosior Engelmann Proc. Amer. Acad. Arts 3: 307. 1856
Synonyms: Opuntia spinosior (Engelmann) Toumey Opuntia spinosior var. neomexicana Toumey
Treatment appears in FNA Volume 4. Mentioned on page 105, 106, 107, 109.
FNA>Volume Importer
 
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{{Treatment/ID
 
{{Treatment/ID
 
|accepted_name=Cylindropuntia spinosior
 
|accepted_name=Cylindropuntia spinosior
|accepted_authority=(Engelmann) F. M. Knuth in C. Backeberg and F. M. Knuth
+
|accepted_authority=(Engelmann) F. M. Knuth
 
|publications={{Treatment/Publication
 
|publications={{Treatment/Publication
 
|title=in C. Backeberg and F. M. Knuth, Kaktus-ABC,
 
|title=in C. Backeberg and F. M. Knuth, Kaktus-ABC,
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}}
 
}}
 
|common_names=Cane cholla;walkingstick cactus
 
|common_names=Cane cholla;walkingstick cactus
|basionyms={{Treatment/ID/Synonym
+
|basionyms={{Treatment/ID/Basionym
 
|name=Opuntia whipplei var. spinosior
 
|name=Opuntia whipplei var. spinosior
 
|authority=Engelmann
 
|authority=Engelmann
 +
|rank=variety
 +
|publication_title=Proc. Amer. Acad. Arts
 +
|publication_place=3: 307. 1856
 
}}
 
}}
 
|synonyms={{Treatment/ID/Synonym
 
|synonyms={{Treatment/ID/Synonym
 
|name=Opuntia spinosior
 
|name=Opuntia spinosior
 
|authority=(Engelmann) Toumey
 
|authority=(Engelmann) Toumey
}}{{Treatment/ID/Synonym
+
|rank=species
 +
}} {{Treatment/ID/Synonym
 
|name=Opuntia spinosior var. neomexicana
 
|name=Opuntia spinosior var. neomexicana
 
|authority=Toumey
 
|authority=Toumey
 +
|rank=variety
 
}}
 
}}
 
|hierarchy=Cactaceae;Cactaceae subfam. Opuntioideae;Cylindropuntia;Cylindropuntia spinosior
 
|hierarchy=Cactaceae;Cactaceae subfam. Opuntioideae;Cylindropuntia;Cylindropuntia spinosior
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}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="tree architecture or arrangement;tree architecture;tree some measurement"><b>Trees,</b> compact, widely branching, 0.4–2 m.</span> <span class="statement" id="st-d0_s1" data-properties="stem segment fixation;stem segment arrangement;stem segment arrangement;stem segment coloration;stem segment coloration;stem segment coloration;stem segment length;stem segment width"><b>Stem </b>segments firmly attached, whorled or subwhorled, green to purple, 5–23 × 1.3–3.5 cm;</span> <span class="statement" id="st-d0_s2" data-properties="tubercle arrangement;tubercle prominence;tubercle shape;tubercle shape;tubercle shape;tubercle atypical some measurement;tubercle some measurement">tubercles crowded, pronounced, oval to narrowly oval, 0.5–1.2 (–1.5) cm;</span> <span class="statement" id="st-d0_s3" data-properties="areole shape;areole shape;areole shape;areole length;areole width">areoles broadly obdeltate to elliptic, 4.5–7 × 2–4 mm;</span> <span class="statement" id="st-d0_s4" data-properties="wool coloration;wool coloration;wool coloration;wool coloration;wool coloration;wool coloration;wool coloration">wool yellow to tan, aging gray to black.</span> <span class="statement" id="st-d0_s5" data-properties="spine atypical count;spine atypical count;spine count;0-areole arrangement;0-areole coloration;0-areole coloration;0-areole coloration;0-areole coloration;areole arrangement"><b>Spines </b>(4–) 6–18 (–24) per areole, at most areoles, interlacing with spines of adjacent areoles, pale tan (rarely yellowish), pinkish to redbrown;</span> <span class="statement" id="st-d0_s6" data-properties="abaxial spine orientation;abaxial spine orientation;abaxial spine orientation;abaxial spine shape;abaxial spine shape;abaxial spine some measurement">abaxial spines erect to usually deflexed, terete, often flattened basally, 8–19 mm;</span> <span class="statement" id="st-d0_s7" data-properties="adaxial spine orientation;adaxial spine orientation;adaxial spine shape">adaxial spines erect or spreading, subterete;</span> <span class="statement" id="st-d0_s8" data-properties="spine architecture">± bristlelike spines at areole abaxial margins;</span> <span class="statement" id="st-d0_s9" data-properties="sheath duration;sheath coloration">sheaths persisting, uniformly whitish, not baggy.</span> <span class="statement" id="st-d0_s10" data-properties="glochid coloration;glochid coloration;glochid coloration;glochid life cycle;glochid coloration;glochid some measurement;adaxial tuft prominence"><b>Glochids </b>in inconspicuous adaxial tuft, yellow to tan, aging gray, 1–2 mm.</span> <span class="statement" id="st-d0_s11" data-properties="inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal shape;inner tepal some measurement;inner tepal architecture or shape"><b>Flowers:</b> inner tepals rose to red-purple, bronze-purple, or yellow, sometimes pale greenish yellow or whitish, spatulate, 18–35 mm, emarginate-apiculate;</span> <span class="statement" id="st-d0_s12" data-properties="filament depth;filament density or coloration;filament coloration">filaments deep purple to pink-purple, sometimes green;</span> <span class="statement" id="st-d0_s13" data-properties="anther coloration">anthers pale-yellow;</span> <span class="statement" id="st-d0_s14" data-properties="style coloration;style coloration;style coloration;style coloration">style white or pink to purple distally;</span> <span class="statement" id="st-d0_s15" data-properties="stigma lobe coloration;stigma lobe coloration;stigma lobe coloration">stigma lobes white to cream.</span> <span class="statement" id="st-d0_s16" data-properties="fruit life cycle;fruit coloration;fruit coloration;fruit coloration;fruit coloration;fruit shape;fruit length;fruit width;fruit texture;fruit relief;fruit architecture"><b>Fruits </b>rarely proliferating, yellow, sometimes tinged reddish to purplish, broadly cylindric, 20–50 × 17–30 mm, fleshy, strongly tuberculate, spineless;</span> <span class="statement" id="st-d0_s17" data-properties="areole atypical count;areole count">areoles 28–50 (–62);</span> <span class="statement" id="st-d0_s18" data-properties="tubercle length or size">tubercles longer in distal portion of fruit;</span> <span class="statement" id="st-d0_s19" data-properties="umbilicus some measurement;umbilicus depth">umbilicus to 10 mm deep.</span> <span class="statement" id="st-d0_s20" data-properties="seed coloration;seed shape;seed shape;seed shape;seed shape;seed shape;seed shape;seed length;seed width;depression count;depression size"><b>Seeds </b>pale-yellow, suborbicular to oval in outline, flattened to warped, 4–5 × 3–4 mm, sides with 0–3 large depressions;</span> <span class="statement" id="st-d0_s21" data-properties="">girdle smooth.</span> <span class="statement" id="st-d0_s22" data-properties="girdle architecture or pubescence or relief;2n chromosome count">2n = 22.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Trees,</b> compact, widely branching, 0.4–2 m. <b>Stem</b> segments firmly attached, whorled or subwhorled, green to purple, 5–23 × 1.3–3.5 cm; tubercles crowded, pronounced, oval to narrowly oval, 0.5–1.2(–1.5) cm; areoles broadly obdeltate to elliptic, 4.5–7 × 2–4 mm; wool yellow to tan, aging gray to black. <b>Spines</b> (4–)6–18(–24) per areole, at most areoles, interlacing with spines of adjacent areoles, pale tan (rarely yellowish), pinkish to red-brown; abaxial spines erect to usually deflexed, terete, often flattened basally, 8–19 mm; adaxial spines erect or spreading, subterete; ± bristlelike spines at areole abaxial margins; sheaths persisting, uniformly whitish, not baggy. <b>Glochids</b> in inconspicuous adaxial tuft, yellow to tan, aging gray, 1–2 mm. <b>Flowers</b>: inner tepals rose to red-purple, bronze-purple, or yellow, sometimes pale greenish yellow or whitish, spatulate, 18–35 mm, emarginate-apiculate; filaments deep purple to pink-purple, sometimes green; anthers pale yellow; style white or pink to purple distally; stigma lobes white to cream. <b>Fruits</b> rarely proliferating, yellow, sometimes tinged reddish to purplish, broadly cylindric, 20–50 × 17–30 mm, fleshy, strongly tuberculate, spineless; areoles 28–50(–62); tubercles longer in distal portion of fruit; umbilicus to 10 mm deep. <b>Seeds</b> pale yellow, suborbicular to oval in outline, flattened to warped, 4–5 × 3–4 mm, sides with 0–3 large depressions; girdle smooth. <b>2n</b> = 22.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
 +
|phenology=Flowering spring–early summer (Apr–Aug).
 
|habitat=Desert and plains grasslands, extending onto Sonoran Desert flats, sandy to loamy soils
 
|habitat=Desert and plains grasslands, extending onto Sonoran Desert flats, sandy to loamy soils
 
|elevation=300-2000 m
 
|elevation=300-2000 m
 
|distribution=Ariz.;N.Mex.;Mexico (Chihuahua;Sonora).
 
|distribution=Ariz.;N.Mex.;Mexico (Chihuahua;Sonora).
|discussion=<p>Cylindropuntia spinosior forms hybrids with C. acanthocarpa var. major, C. arbuscula, C. fulgida (see 6. C. ×kelvinensis), C. leptocaulis (see discussion under 3. C. ×tetracantha), and C. versicolor (= C. ×grantiorum P. V. Heath). Introgression between C. spinosior and C. imbricata occurs in a more or less continuous band from central New Mexico to Chihuahua, Mexico. The hybrids between C. spinosior and C. acanthocarpa var. major are sprawling shrubs with irregular branching pattern and have spine clusters with one or more spines longer than others and fleshy fruits, some with one to few spines per fruit. Chromosome numbers of hybrids are all reported as 2n = 22.</p>
+
|discussion=<p><i>Cylindropuntia spinosior</i> forms hybrids with <i>C. acanthocarpa </i>var.<i> major</i>, <i>C. arbuscula</i>, <i>C. fulgida</i> (see 6. <i>C. ×kelvinensis</i>), <i>C. leptocaulis</i> (see discussion under 3. <i>C. ×tetracantha</i>), and <i>C. versicolor</i> (= C. ×grantiorum P. V. Heath). Introgression between <i>C. spinosior</i> and <i>C. imbricata</i> occurs in a more or less continuous band from central New Mexico to Chihuahua, Mexico. The hybrids between <i>C. spinosior</i> and <i>C. acanthocarpa </i>var.<i> major</i> are sprawling shrubs with irregular branching pattern and have spine clusters with one or more spines longer than others and fleshy fruits, some with one to few spines per fruit. Chromosome numbers of hybrids are all reported as 2n = 22.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Cylindropuntia spinosior
 
name=Cylindropuntia spinosior
|author=
+
|authority=(Engelmann) F. M. Knuth
|authority=(Engelmann) F. M. Knuth in C. Backeberg and F. M. Knuth
 
 
|rank=species
 
|rank=species
 
|parent rank=genus
 
|parent rank=genus
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|basionyms=Opuntia whipplei var. spinosior
 
|basionyms=Opuntia whipplei var. spinosior
 
|family=Cactaceae
 
|family=Cactaceae
 +
|phenology=Flowering spring–early summer (Apr–Aug).
 
|habitat=Desert and plains grasslands, extending onto Sonoran Desert flats, sandy to loamy soils
 
|habitat=Desert and plains grasslands, extending onto Sonoran Desert flats, sandy to loamy soils
 
|elevation=300-2000 m
 
|elevation=300-2000 m
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|publication year=1935
 
|publication year=1935
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_211.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V4/V4_211.xml
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|genus=Cylindropuntia
 
|genus=Cylindropuntia
 
|species=Cylindropuntia spinosior
 
|species=Cylindropuntia spinosior
|0-areole arrangement=interlacing
 
|0-areole coloration=pinkish;redbrown
 
|2n chromosome count=22
 
|abaxial spine orientation=erect;usually deflexed
 
|abaxial spine shape=flattened;terete
 
|abaxial spine some measurement=8mm;19mm
 
|adaxial spine orientation=spreading;erect
 
|adaxial spine shape=subterete
 
|adaxial tuft prominence=inconspicuous
 
|anther coloration=pale-yellow
 
|areole arrangement=adjacent
 
|areole atypical count=50;62
 
|areole count=28;50
 
|areole length=4.5mm;7mm
 
|areole shape=broadly obdeltate;elliptic
 
|areole width=2mm;4mm
 
|depression count=0;3
 
|depression size=large
 
|filament coloration=green
 
|filament density or coloration=purple to pink-purple
 
|filament depth=deep
 
|fruit architecture=spineless
 
|fruit coloration=tinged reddish;purplish
 
|fruit length=20mm;50mm
 
|fruit life cycle=proliferating
 
|fruit relief=tuberculate
 
|fruit shape=cylindric
 
|fruit texture=fleshy
 
|fruit width=17mm;30mm
 
|girdle architecture or pubescence or relief=smooth
 
|glochid coloration=gray;yellow;tan
 
|glochid life cycle=aging
 
|glochid some measurement=1mm;2mm
 
|inner tepal architecture or shape=emarginate-apiculate
 
|inner tepal coloration=whitish;yellow;pale greenish;yellow;bronze-purple;yellow;bronze-purple;rose to red-purple
 
|inner tepal shape=spatulate
 
|inner tepal some measurement=18mm;35mm
 
|seed coloration=pale-yellow
 
|seed length=4mm;5mm
 
|seed shape=flattened;warped
 
|seed width=3mm;4mm
 
|sheath coloration=whitish
 
|sheath duration=persisting
 
|spine architecture=bristlelike
 
|spine atypical count=18;24
 
|spine count=6;18
 
|stem segment arrangement=subwhorled;whorled
 
|stem segment coloration=green;purple
 
|stem segment fixation=attached
 
|stem segment length=5cm;23cm
 
|stem segment width=1.3cm;3.5cm
 
|stigma lobe coloration=white;cream
 
|style coloration=pink;purple
 
|tree architecture=branching
 
|tree architecture or arrangement=compact
 
|tree some measurement=0.4m;2m
 
|tubercle arrangement=crowded
 
|tubercle atypical some measurement=1.2cm;1.5cm
 
|tubercle length or size=longer
 
|tubercle prominence=pronounced
 
|tubercle shape=oval;narrowly oval
 
|tubercle some measurement=0.5cm;1.2cm
 
|umbilicus depth=deep
 
|umbilicus some measurement=0mm;10mm
 
|wool coloration=gray;black
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Cylindropuntia]]
 
-->[[Category:Treatment]][[Category:Cylindropuntia]]

Latest revision as of 22:57, 5 November 2020

Trees, compact, widely branching, 0.4–2 m. Stem segments firmly attached, whorled or subwhorled, green to purple, 5–23 × 1.3–3.5 cm; tubercles crowded, pronounced, oval to narrowly oval, 0.5–1.2(–1.5) cm; areoles broadly obdeltate to elliptic, 4.5–7 × 2–4 mm; wool yellow to tan, aging gray to black. Spines (4–)6–18(–24) per areole, at most areoles, interlacing with spines of adjacent areoles, pale tan (rarely yellowish), pinkish to red-brown; abaxial spines erect to usually deflexed, terete, often flattened basally, 8–19 mm; adaxial spines erect or spreading, subterete; ± bristlelike spines at areole abaxial margins; sheaths persisting, uniformly whitish, not baggy. Glochids in inconspicuous adaxial tuft, yellow to tan, aging gray, 1–2 mm. Flowers: inner tepals rose to red-purple, bronze-purple, or yellow, sometimes pale greenish yellow or whitish, spatulate, 18–35 mm, emarginate-apiculate; filaments deep purple to pink-purple, sometimes green; anthers pale yellow; style white or pink to purple distally; stigma lobes white to cream. Fruits rarely proliferating, yellow, sometimes tinged reddish to purplish, broadly cylindric, 20–50 × 17–30 mm, fleshy, strongly tuberculate, spineless; areoles 28–50(–62); tubercles longer in distal portion of fruit; umbilicus to 10 mm deep. Seeds pale yellow, suborbicular to oval in outline, flattened to warped, 4–5 × 3–4 mm, sides with 0–3 large depressions; girdle smooth. 2n = 22.


Phenology: Flowering spring–early summer (Apr–Aug).
Habitat: Desert and plains grasslands, extending onto Sonoran Desert flats, sandy to loamy soils
Elevation: 300-2000 m

Distribution

V4 211-distribution-map.gif

Ariz., N.Mex., Mexico (Chihuahua, Sonora).

Discussion

Cylindropuntia spinosior forms hybrids with C. acanthocarpa var. major, C. arbuscula, C. fulgida (see 6. C. ×kelvinensis), C. leptocaulis (see discussion under 3. C. ×tetracantha), and C. versicolor (= C. ×grantiorum P. V. Heath). Introgression between C. spinosior and C. imbricata occurs in a more or less continuous band from central New Mexico to Chihuahua, Mexico. The hybrids between C. spinosior and C. acanthocarpa var. major are sprawling shrubs with irregular branching pattern and have spine clusters with one or more spines longer than others and fleshy fruits, some with one to few spines per fruit. Chromosome numbers of hybrids are all reported as 2n = 22.

Selected References

None.

Lower Taxa

None.
... more about "Cylindropuntia spinosior"
Donald J. Pinkava +
(Engelmann) F. M. Knuth in C. Backeberg and F. M. Knuth +
Opuntia whipplei var. spinosior +
Cane cholla +  and walkingstick cactus +
Ariz. +, N.Mex. +, Mexico (Chihuahua +  and Sonora). +
300-2000 m +
Desert and plains grasslands, extending onto Sonoran Desert flats, sandy to loamy soils +
in C. Backeberg and F. M. Knuth, Kaktus-ABC, +
Opuntia spinosior +  and Opuntia spinosior var. neomexicana +
Cylindropuntia spinosior +
Cylindropuntia +
species +