Carpinus

Trees, 8–25 m; trunks usually 1, branching mostly deliquescent, trunk and branches irregularly longitudinally ridged, fluted. Bark of trunk and branches bluish to brownish gray, thin, smooth, close [thicker, broken or shredded]; lenticels generally inconspicuous. Wood nearly white to light brown, very hard and heavy, texture fine. Branches, branchlets, and twigs conspicuously 2-ranked; young twigs differentiated into long and short shoots. Winter buds sessile, ovoid, 4-angled in cross section, apex acute; scales many, imbricate, smooth. Leaves on long and short shoots, 2-ranked. Leaf blade narrowly ovate to ovate, elliptic, or obovate with 10 or more pairs of lateral veins, 3–12 × 3–6 cm, thin, margins doubly serrate to serrulate; surfaces abaxially glabrous to tomentose, sometimes covered with small glands. Inflorescences: staminate catkins solitary or in small racemose clusters, lateral, formed previous growing season and enclosed [exposed] in buds during winter, expanding with leaves; pistillate catkins distal to staminate on short, leafy new growth, solitary, ± erect, elongate; bracts and flowers uncrowded. Staminate flowers in catkins 3 per scale, crowded together on pilose receptacle; stamens 3(–6), short; filaments often distinct part way to base; anthers divided into 2 parts, each 1-locular, apex pilose, Pistillate flowers 2 per bract. Infructescences loose racemose clusters of paired bracts, clusters pendulous, elongate; paired bracts deciduous with fruit, expanded, (1–)3-lobed, variously toothed, foliaceous, each bract subtending 1 fruit. Fruits small nutlets, deltoid, longitudinally ribbed, often crowned with persistent sepals and styles. x = 8.

Distribution

Mostly north temperate zone, Europe, Asia (s to India, Iran).

Discussion

Species ca. 25 (1 in the flora).

In the flora, Carpinus consists of a single native species, C. caroliniana, which is composed of two fairly distinctive geographic races (J. J. Furlow 1987, 1987b), treated here as subspecies. Worldwide it includes about 25 species, some of which become large trees. The European C. betulus is frequently planted in North America and persists long after other signs of human development have vanished. It seldom escapes, however, and it has not become naturalized. In the mountains of Mexico and Central America, the larger C. tropicalis (Donnell Smith) Lundell is widespread in the temperate forest zone.

Closely related to Ostrya, Carpinus is easily recognized by its smooth, gray, often fluted stems and racemose infructescences consisting of pairs of uncrowded, foliaceous, 3-lobed bracts, each subtending a small triangular nutlet. The staminate (but not the pistillate) catkins develop in the autumn and are enclosed within buds throughout the winter prior to anthesis (in Ostrya, these are exposed during the winter). The pistillate catkins are produced on the first new growth in the spring.

Of relatively minor economic importance, Carpinus has limited use for its very hard wood, especially in Europe, where it is used for making mallet heads, tool handles, levers, and other small, hard, wooden objects.

Selected References

Furlow, J. J. 1987. The Carpinus caroliniana complex in North America. I. A multivariate analysis of geographical variation. Syst. Bot. 12: 21--40.

Furlow, J. J. 1987b. The Carpinus caroliniana complex in North America. II. Systematics. Syst. Bot. 12: 416--434.

Winkler, H. 1914. Neue Revision der Gattung Carpinus. Bot. Jahrb. Syst. 15(suppl.): 488--508.

Winstead, J. E., B. J. Smith, and G. I. Wardell. 1977. Fruit weight clines in populations of ash, ironwood, cherry, dogwood and maple. Castanea 42: 56--60.

Lower Taxa

Carpinus caroliniana