Difference between revisions of "Opuntia polyacantha"

Haworth

Suppl. Pl. Succ., 82. 1819.

Common names: Starvation pricklypear
Basionym: Cactus ferox Nuttall
Synonyms: Tunas polyacantha (Haworth) Nieuwland & Lunell
Treatment appears in FNA Volume 4. Mentioned on page 124, 125, 132, 133.
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--><span class="statement" id="st-d0_s0" data-properties="shrub position;shrub some measurement;branch growth form or orientation"><b>Shrubs,</b> low, 10–25 cm, with ± prostrate branches.</span> <span class="statement" id="st-d0_s1" data-properties="stem segment fusion;stem segment coloration;stem segment shape;stem segment shape;stem segment shape;stem segment shape;stem segment length;stem segment width;stem segment position;stem segment relief"><b>Stem </b>segments not easily detached, green, elliptic to narrowly to broadly obovate to circular, 4–27 × 2–18 cm, low tuberculate;</span> <span class="statement" id="st-d0_s2" data-properties="areole count;areole shape;areole some measurement">areoles 4–14 per diagonal row across midstem segment, subcircular, 3–6 mm;</span> <span class="statement" id="st-d0_s3" data-properties="wool coloration;wool coloration;wool coloration">wool tan to brown.</span> <span class="statement" id="st-d0_s4" data-properties="spine shape;spine shape;spine shape;spine fragility or size;spine shape;spine architecture;spine course;spine course;spine course;areole position or shape"><b>Spines </b>at all or only distal areoles of stem segment, terete to flattened, stout to acicular to bristlelike, straight to curling, of 1 or 2 kinds;</span> <span class="statement" id="st-d0_s5" data-properties="whole_organism count;whole_organism count;whole_organism orientation;whole_organism course;whole_organism course;whole_organism orientation;whole_organism orientation;whole_organism orientation;whole_organism orientation;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism coloration;whole_organism length;whole_organism atypical some measurement;whole_organism atypical some measurement;whole_organism some measurement;direction variability">if 1 kind: 0–18 per areole, spreading and curling in various directions, sometimes straight, erect, ascending to deflexed, yellow to dark-brown to black, turning gray, pink-gray to gray-brown, longest (35–) 40–90 (–185) mm;</span> <span class="statement" id="st-d0_s6" data-properties="whole_organism count">if ± 2 kinds: major spines (0–) 1–5, reflexed to porrect, yellowbrown to brown to gray, longest 20–150 mm;</span> <span class="statement" id="st-d0_s7" data-properties="spine size;spine atypical count;spine count;spine orientation;spine orientation;spine orientation;spine coloration;spine coloration;spine coloration;spine coloration;spine length;spine some measurement;minor spine atypical count;minor spine count;minor spine orientation;minor spine coloration;minor spine coloration;minor spine coloration;minor spine length;minor spine some measurement">minor spines (0–) 5–11, deflexed, white to white-gray, longest 4–16 mm.</span> <span class="statement" id="st-d0_s8" data-properties="glochid prominence;glochid coloration;glochid coloration;glochid coloration;glochid life cycle;glochid coloration;glochid some measurement;tuft width;tuft architecture or shape;tuft shape"><b>Glochids </b>inconspicuous, in narrow, tidy crescent at adaxial edge of areole or in broad, brushy crescent and tuft, yellow to reddish, aging brown, to 10 mm.</span> <span class="statement" id="st-d0_s9" data-properties="inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal some measurement"><b>Flowers:</b> inner tepals yellow to magenta throughout, 25–40 mm;</span> <span class="statement" id="st-d0_s10" data-properties="filament coloration;filament coloration;filament coloration;filament coloration;filament coloration;filament coloration;filament coloration;filament coloration;filament coloration">filaments white, yellow, or red to magenta (flowers may superficially appear bicolored);</span> <span class="statement" id="st-d0_s11" data-properties="anther coloration">anthers yellow;</span> <span class="statement" id="st-d0_s12" data-properties="style coloration;style coloration;style coloration">style white to pale-pink;</span> <span class="statement" id="st-d0_s13" data-properties="stigma lobe coloration">stigma lobes green.</span> <span class="statement" id="st-d0_s14" data-properties="fruit coloration;fruit coloration;fruit coloration;fruit shape;fruit length;fruit width;fruit condition or texture;fruit pubescence;fruit architecture or shape"><b>Fruits </b>tan to brown, ± cylindric, 15–45 × 12–25 mm, dry at maturity, glabrous, sometimes burlike;</span> <span class="statement" id="st-d0_s15" data-properties="areole count;areole position or shape;areole some measurement;spine count">areoles 10–33, each or only distal areoles bearing 3–16 spines, 4–20 mm.</span> <span class="statement" id="st-d0_s16" data-properties="seed coloration;seed coloration;seed coloration;seed shape;seed shape;seed shape;seed shape;seed shape;seed length;seed width"><b>Seeds </b>tan to gray, flattened, warped, oblong to subcircular, 3–7 × 2–4 mm;</span> <span class="statement" id="st-d0_s17" data-properties="girdle prominence;girdle some measurement">girdle protruding 1–2 mm.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Shrubs,</b> low, 10–25 cm, with ± prostrate branches. <b>Stem</b> segments not easily detached, green, elliptic to narrowly to broadly obovate to circular, 4–27 × 2–18 cm, low tuberculate; areoles 4–14 per diagonal row across midstem segment, subcircular, 3–6 mm; wool tan to brown. <b>Spines</b> at all or only distal areoles of stem segment, terete to flattened, stout to acicular to bristlelike, straight to curling, of 1 or 2 kinds; if 1 kind: 0–18 per areole, spreading and curling in various directions, sometimes straight, erect, ascending to deflexed, yellow to dark brown to black, turning gray, pink-gray to gray-brown, longest (35–)40–90(–185) mm; if ± 2 kinds: major spines (0–)1–5, reflexed to porrect, yellow-brown to brown to gray, longest 20–150 mm; minor spines (0–)5–11, deflexed, white to white-gray, longest 4–16 mm. <b>Glochids</b> inconspicuous, in narrow, tidy crescent at adaxial edge of areole or in broad, brushy crescent and tuft, yellow to reddish, aging brown, to 10 mm. <b>Flowers</b>: inner tepals yellow to magenta throughout, 25–40 mm; filaments white, yellow, or red to magenta (flowers may superficially appear bicolored); anthers yellow; style white to pale pink; stigma lobes green. <b>Fruits</b> tan to brown, ± cylindric, 15–45 × 12–25 mm, dry at maturity, glabrous, sometimes burlike; areoles 10–33, each or only distal areoles bearing 3–16 spines, 4–20 mm. <b>Seeds</b> tan to gray, flattened, warped, oblong to subcircular, 3–7 × 2–4 mm; girdle protruding 1–2 mm.</span><!--
  
 
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|publication year=1819
 
|publication year=1819
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_283.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V4/V4_283.xml
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|subfamily=Cactaceae subfam. Opuntioideae
 
|genus=Opuntia
 
|genus=Opuntia
 
|species=Opuntia polyacantha
 
|species=Opuntia polyacantha
|anther coloration=yellow
 
|areole count=10;33
 
|areole position or shape=distal;distal
 
|areole shape=subcircular
 
|areole some measurement=4mm;20mm
 
|branch growth form or orientation=prostrate
 
|direction variability=various
 
|filament coloration=red;magenta
 
|fruit architecture or shape=burlike
 
|fruit coloration=tan;brown
 
|fruit condition or texture=dry
 
|fruit length=15mm;45mm
 
|fruit pubescence=glabrous
 
|fruit shape=cylindric
 
|fruit width=12mm;25mm
 
|girdle prominence=protruding
 
|girdle some measurement=1mm;2mm
 
|glochid coloration=brown;yellow;reddish
 
|glochid life cycle=aging
 
|glochid prominence=inconspicuous
 
|glochid some measurement=0mm;10mm
 
|inner tepal coloration=yellow;magenta
 
|inner tepal some measurement=25mm;40mm
 
|minor spine atypical count=0;5
 
|minor spine coloration=white;white-gray
 
|minor spine count=5;11
 
|minor spine length=longest
 
|minor spine orientation=deflexed
 
|minor spine some measurement=4mm;16mm
 
|seed coloration=tan;gray
 
|seed length=3mm;7mm
 
|seed shape=oblong;subcircular
 
|seed width=2mm;4mm
 
|shrub position=low
 
|shrub some measurement=10cm;25cm
 
|spine architecture=bristlelike
 
|spine atypical count=0;1
 
|spine coloration=yellowbrown;brown
 
|spine count=3;16
 
|spine course=straight;curling
 
|spine fragility or size=stout
 
|spine length=longest
 
|spine orientation=reflexed;porrect
 
|spine shape=acicular;terete;flattened
 
|spine size=major
 
|spine some measurement=20mm;150mm
 
|stem segment coloration=green
 
|stem segment fusion=detached
 
|stem segment length=4cm;27cm
 
|stem segment position=low
 
|stem segment relief=tuberculate
 
|stem segment shape=elliptic;to narrowly broadly obovate
 
|stem segment width=2cm;18cm
 
|stigma lobe coloration=green
 
|style coloration=white;pale-pink
 
|tuft architecture or shape=brushy
 
|tuft shape=crescent
 
|tuft width=broad
 
|whole_organism atypical some measurement=90mm;185mm
 
|whole_organism coloration=pink-gray;gray-brown
 
|whole_organism count=2;0;18
 
|whole_organism course=straight;curling
 
|whole_organism length=longest
 
|whole_organism orientation=ascending;deflexed
 
|whole_organism some measurement=40mm;90mm
 
|wool coloration=tan;brown
 
 
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Revision as of 14:03, 27 July 2019

Shrubs, low, 10–25 cm, with ± prostrate branches. Stem segments not easily detached, green, elliptic to narrowly to broadly obovate to circular, 4–27 × 2–18 cm, low tuberculate; areoles 4–14 per diagonal row across midstem segment, subcircular, 3–6 mm; wool tan to brown. Spines at all or only distal areoles of stem segment, terete to flattened, stout to acicular to bristlelike, straight to curling, of 1 or 2 kinds; if 1 kind: 0–18 per areole, spreading and curling in various directions, sometimes straight, erect, ascending to deflexed, yellow to dark brown to black, turning gray, pink-gray to gray-brown, longest (35–)40–90(–185) mm; if ± 2 kinds: major spines (0–)1–5, reflexed to porrect, yellow-brown to brown to gray, longest 20–150 mm; minor spines (0–)5–11, deflexed, white to white-gray, longest 4–16 mm. Glochids inconspicuous, in narrow, tidy crescent at adaxial edge of areole or in broad, brushy crescent and tuft, yellow to reddish, aging brown, to 10 mm. Flowers: inner tepals yellow to magenta throughout, 25–40 mm; filaments white, yellow, or red to magenta (flowers may superficially appear bicolored); anthers yellow; style white to pale pink; stigma lobes green. Fruits tan to brown, ± cylindric, 15–45 × 12–25 mm, dry at maturity, glabrous, sometimes burlike; areoles 10–33, each or only distal areoles bearing 3–16 spines, 4–20 mm. Seeds tan to gray, flattened, warped, oblong to subcircular, 3–7 × 2–4 mm; girdle protruding 1–2 mm.

Discussion

Varieties 5 (5 in the flora).

Populations of Opuntia polyacantha with spines few or absent (especially var. hystricina) were the basis for several names including O. juniperina, O. utahensis, and O. rhodantha.

Selected References

None.

Key

1 Areoles 4-6 per diagonal row across midstem segment; stem segments 4-7 × 2-3(-5) cm, elliptic to very narrowly obovate; Texas, New Mexico, Mexico Opuntia polyacantha var. arenaria
1 Areoles 6-14 per diagonal row across midstem segment; stem segments 6-27 × 5-18 cm, obovate to circular or, if elliptic, longer than 9 cm; widespread in w United States and Canada > 2
2 Spines of 2 kinds: major spines 1-3 per areole, usually deflexed on terminal stem segment, but sometimes ascending at stem segment apex; minor spines 1-3 per areole, deflexed, white, subtending major spines > 3
2 Spines of 1 kind, grading in length, from ascending to deflexed, 7-18 per areole, longest spines of terminal stem segment porrect to ascending, shorter spines deflexed to porrect > 4
3 Longest spines 2-4 cm; areoles 6-13 mm apart; stem segments 6-12 cm; Great Plains, n and e Great Basin Desert Opuntia polyacantha var. polyacantha
3 Longest spines 4.5-12 cm; areoles 12-30 mm apart; stem segments 10-27 cm; Arizona and Utah border Opuntia polyacantha var. nicholii
4 Spines usually dark brown, those of proximal stem segments like those of terminal stem segments, porrect to ascending, straight and stiff; fruit areoles 11-21, spiny but not burlike Opuntia polyacantha var. hystricina
4 Spines usually pale, those of proximal stem segments not like those of terminals stem segments, more numerous, longer and more reflexed, straight or flexuous and/or hairlike or threadlike; fruit areoles 20-33, spiny, forming dense bur Opuntia polyacantha var. erinacea