Difference between revisions of "Cyrtomium"

C. Presl

Tent. Pterid. 86. 1836.

Common names: Asiatic holly fern
Etymology: Greek cyrtoma, arch, for the arched veins
Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties="plant growth form or habitat;plant growth form or habitat"><b>Plants </b>terrestrial or on rock.</span> <span class="statement" id="st-d0_s1" data-properties="stem orientation;stem orientation;stolon count"><b>Stems </b>erect or ascending, stolons absent.</span> <span class="statement" id="st-d0_s2" data-properties="leaf architecture;leaf duration"><b>Leaves </b>monomorphic, evergreen.</span> <span class="statement" id="st-d0_s3" data-properties="petiole length;base shape"><b>Petiole </b>± 1/2–3/4 length of blade, base not swollen;</span> <span class="statement" id="st-d0_s4" data-properties="vascular-bundle count;vascular-bundle arrangement;vascular-bundle shape">vascular-bundles more than 3, arranged in arc, ± round in cross-section.</span> <span class="statement" id="st-d0_s5" data-properties="blade shape;blade architecture or shape;pinna shape;distal pinna size;blade texture;blade texture;pinna shape"><b>Blade </b>oblong-lanceolate, 1-pinnate with pinnae not lobed, distal pinnae only slightly smaller, blade ending in basally lobed pinna ± similar to lateral pinnae, papery or somewhat leathery.</span> <span class="statement" id="st-d0_s6" data-properties="pinna architecture;segment margin shape;segment margin architecture or shape"><b>Pinnae </b>not articulate to rachis, segment margins crenate to spinulose, sometimes also coarsely dentate;</span> <span class="statement" id="st-d0_s7" data-properties="proximal pinna size;proximal pinna architecture;proximal pinna architecture or shape;proximal pinna shape;base orientation;base development">proximal pinnae not reduced, petiolulate, equilateral or somewhat inequilateral with acroscopic base more developed;</span> <span class="statement" id="st-d0_s8" data-properties="costa architecture;groove architecture">costae adaxially grooved, grooves continuous from rachis to costae;</span> <span class="statement" id="st-d0_s9" data-properties="indument count;scale shape">indument of filiform scales abaxially, absent adaxially.</span> <span class="statement" id="st-d0_s10" data-properties="vein architecture;included veinlet count"><b>Veins </b>elaborately anastomosing, areoles formed with 1–3 included veinlets.</span> <span class="statement" id="st-d0_s11" data-properties="between midrib and margin sorus shape"><b>Sori </b>in 2 or more rows between midrib and margin, round.</span> <span class="statement" id="st-d0_s12" data-properties="indusium architecture;indusium duration;indusium duration"><b>Indusia </b>peltate, persistent or caducous.</span> <span class="statement" id="st-d0_s13" data-properties="fold shape"><b>Spores </b>brown, with inflated folds or wings.</span> <span class="statement" id="st-d0_s14" data-properties="spore coloration;x chromosome count">x = 41.</span><!--
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--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>terrestrial or on rock. <b>Stems</b> erect or ascending, stolons absent. <b>Leaves</b> monomorphic, evergreen. <b>Petiole</b> ± 1/2–3/4 length of blade, base not swollen; vascular bundles more than 3, arranged in arc, ± round in cross section. <b>Blade</b> oblong-lanceolate, 1-pinnate with pinnae not lobed, distal pinnae only slightly smaller, blade ending in basally lobed pinna ± similar to lateral pinnae, papery or somewhat leathery. <b>Pinnae</b> not articulate to rachis, segment margins crenate to spinulose, sometimes also coarsely dentate; proximal pinnae not reduced, petiolulate, equilateral or somewhat inequilateral with acroscopic base more developed; costae adaxially grooved, grooves continuous from rachis to costae; indument of filiform scales abaxially, absent adaxially. <b>Veins</b> elaborately anastomosing, areoles formed with 1–3 included veinlets. <b>Sori</b> in 2 or more rows between midrib and margin, round. <b>Indusia</b> peltate, persistent or caducous. <b>Spores</b> brown, with inflated folds or wings. <b>x</b> = 41.</span><!--
  
 
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|publication year=1836
 
|publication year=1836
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_578.xml
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_578.xml
 
|genus=Cyrtomium
 
|genus=Cyrtomium
|base development=developed
 
|base orientation=acroscopic
 
|base shape=not swollen
 
|between midrib and margin sorus shape=round
 
|blade architecture or shape=1-pinnate
 
|blade shape=oblong-lanceolate
 
|blade texture=leathery;papery
 
|costa architecture=grooved
 
|distal pinna size=smaller
 
|fold shape=inflated
 
|groove architecture=continuous
 
|included veinlet count=1;3
 
|indument count=absent
 
|indusium architecture=peltate
 
|indusium duration=caducous;persistent
 
|leaf architecture=monomorphic
 
|leaf duration=evergreen
 
|petiole length=1/2 length of blade;3/4 length of blade
 
|pinna architecture=not articulate
 
|pinna shape=lobed;not lobed
 
|plant growth form or habitat=on rock;terrestrial
 
|proximal pinna architecture=petiolulate
 
|proximal pinna architecture or shape=equilateral
 
|proximal pinna shape=inequilateral
 
|proximal pinna size=not reduced
 
|scale shape=filiform
 
|segment margin architecture or shape=dentate
 
|segment margin shape=crenate to spinulose
 
|spore coloration=brown
 
|stem orientation=ascending;erect
 
|stolon count=absent
 
|vascular-bundle arrangement=arranged
 
|vascular-bundle shape=round
 
|vein architecture=anastomosing
 
|x chromosome count=41
 
 
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Revision as of 13:23, 27 July 2019

Plants terrestrial or on rock. Stems erect or ascending, stolons absent. Leaves monomorphic, evergreen. Petiole ± 1/2–3/4 length of blade, base not swollen; vascular bundles more than 3, arranged in arc, ± round in cross section. Blade oblong-lanceolate, 1-pinnate with pinnae not lobed, distal pinnae only slightly smaller, blade ending in basally lobed pinna ± similar to lateral pinnae, papery or somewhat leathery. Pinnae not articulate to rachis, segment margins crenate to spinulose, sometimes also coarsely dentate; proximal pinnae not reduced, petiolulate, equilateral or somewhat inequilateral with acroscopic base more developed; costae adaxially grooved, grooves continuous from rachis to costae; indument of filiform scales abaxially, absent adaxially. Veins elaborately anastomosing, areoles formed with 1–3 included veinlets. Sori in 2 or more rows between midrib and margin, round. Indusia peltate, persistent or caducous. Spores brown, with inflated folds or wings. x = 41.

Distribution

North America, Asia, Africa including Madagascar, Pacific Islands (Hawaii).

Discussion

This difficult genus requires further systematic study. Estimates of species numbers have ranged from 9 (C. Christensen 1930) to 59 (Shing K. H. 1965). The group might better be considered a subgenus of Polystichum, from which it is poorly differentiated morphologically.

Species ca. 15 (2 in the flora).

Key

1 Pinnae 4-10(-12) pairs, leathery, shiny adaxially, margins sometimes undulate or coarsely dentate, but not minutely crenulate or denticulate. Cyrtomium falcatum
1 Pinnae (8-)10-25 pairs, papery, not shiny adaxially, margins minutely crenulate-denticulate. Cyrtomium fortunei var. fortunei