Difference between revisions of "Woodsia ilvensis"

(Linnaeus) R. Brown

Trans. Linn. Soc. London, Bot. 11: 173. 1813.

Common names: Rusty cliff fern woodsie de l'île d'Elbe
Basionym: Acrostichum ilvense Linnaeus
Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties="stem architecture or arrangement;stem orientation;stem orientation;stem orientation;petiole base count;petiole base duration"><b>Stems </b>compact, erect to ascending, with abundant persistent petiole bases of ± equal length;</span> <span class="statement" id="st-d0_s1" data-properties="scale coloration;scale shape">scales uniformly brown, lanceolate.</span> <span class="statement" id="st-d0_s2" data-properties="leaf length;leaf width"><b>Leaves </b>4.5–25 × 1.2–3.5 cm.</span> <span class="statement" id="st-d0_s3" data-properties="petiole coloration;petiole coloration;petiole architecture;petiole fragility;node shape"><b>Petiole </b>usually brown or dark purple when mature, articulate above base at swollen node, relatively brittle and easily shattered.</span> <span class="statement" id="st-d0_s4" data-properties="blade shape;blade architecture or shape;gland count;gland coating"><b>Blade </b>narrowly lanceolate, usually 2-pinnate proximally, lacking glands, never viscid;</span> <span class="statement" id="st-d0_s5" data-properties="hair count">rachis usually with abundant hairs and scales.</span> <span class="statement" id="st-d0_s6" data-properties="pinna shape;pinna shape;pinna shape;pinna length or size;pinna shape;apex shape;apex shape"><b>Pinnae </b>ovatelanceolate to deltate, longer than wide, abruptly tapered to a rounded or broadly acute apex;</span> <span class="statement" id="st-d0_s7" data-properties="pair count">largest pinnae with 4–9 pairs of pinnules;</span> <span class="statement" id="st-d0_s8" data-properties="scale shape;hair architecture;hair arrangement or density">abaxial surface with mixture of hairs and linear-lanceolate scales, adaxial surface with multicellular hairs concentrated along midrib.</span> <span class="statement" id="st-d0_s9" data-properties="pinnule shape;pinnule shape;pinnule shape"><b>Pinnules </b>entire or crenate, rarely shallowly lobed;</span> <span class="statement" id="st-d0_s10" data-properties="margin reflectance;margin width;margin architecture or pubescence or shape;hair architecture;projection count;projection coloration or reflectance">margins nonlustrous, thin, ciliate with multicellular hairs, lacking translucent projections.</span> <span class="statement" id="st-d0_s11" data-properties="vein tip size;hydathode coloration;hydathode prominence"><b>Vein </b>tips frequently enlarged to form whitish hydathodes visible adaxially.</span> <span class="statement" id="st-d0_s12" data-properties="indusium architecture or arrangement;indusium count;indusium length or size;segment size or width;segment shape;sporangium life cycle"><b>Indusia </b>of narrow, hairlike segments, these uniseriate throughout, composed of cells many times longer than wide, usually surpassing mature sporangia.</span> <span class="statement" id="st-d0_s13" data-properties="spore some measurement;2n chromosome count"><b>Spores </b>averaging 39–46 µm. <b>2n</b> = 82.</span><!--
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--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact, erect to ascending, with abundant persistent petiole bases of ± equal length; scales uniformly brown, lanceolate. <b>Leaves</b> 4.5–25 × 1.2–3.5 cm. <b>Petiole</b> usually brown or dark purple when mature, articulate above base at swollen node, relatively brittle and easily shattered. <b>Blade</b> narrowly lanceolate, usually 2-pinnate proximally, lacking glands, never viscid; rachis usually with abundant hairs and scales. <b>Pinnae</b> ovate-lanceolate to deltate, longer than wide, abruptly tapered to a rounded or broadly acute apex; largest pinnae with 4–9 pairs of pinnules; abaxial surface with mixture of hairs and linear-lanceolate scales, adaxial surface with multicellular hairs concentrated along midrib. <b>Pinnules</b> entire or crenate, rarely shallowly lobed; margins nonlustrous, thin, ciliate with multicellular hairs, lacking translucent projections. <b>Vein</b> tips frequently enlarged to form whitish hydathodes visible adaxially. <b>Indusia</b> of narrow, hairlike segments, these uniseriate throughout, composed of cells many times longer than wide, usually surpassing mature sporangia. <b>Spores</b> averaging 39–46 µm. <b>2n</b> = 82.</span><!--
  
 
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|phenology=Sporulating summer–early fall.
 
|habitat=Cliffs and rocky slopes, found on variety of substrates including serpentine
 
|habitat=Cliffs and rocky slopes, found on variety of substrates including serpentine
 
|elevation=0–1500 m
 
|elevation=0–1500 m
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|basionyms=Acrostichum ilvense
 
|basionyms=Acrostichum ilvense
 
|family=Dryopteridaceae
 
|family=Dryopteridaceae
 +
|phenology=Sporulating summer–early fall.
 
|habitat=Cliffs and rocky slopes, found on variety of substrates including serpentine
 
|habitat=Cliffs and rocky slopes, found on variety of substrates including serpentine
 
|elevation=0–1500 m
 
|elevation=0–1500 m
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|publication year=1813
 
|publication year=1813
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_349.xml
+
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_349.xml
 
|genus=Woodsia
 
|genus=Woodsia
 
|species=Woodsia ilvensis
 
|species=Woodsia ilvensis
|2n chromosome count=82
 
|apex shape=acute;rounded
 
|blade architecture or shape=2-pinnate
 
|blade shape=lanceolate
 
|gland coating=viscid
 
|gland count=lacking
 
|hair architecture=multicellular;multicellular
 
|hair arrangement or density=concentrated
 
|hair count=abundant
 
|hydathode coloration=whitish
 
|hydathode prominence=visible
 
|indusium architecture or arrangement=uniseriate
 
|indusium count=many
 
|indusium length or size=longer than wide
 
|leaf length=4.5cm;25cm
 
|leaf width=1.2cm;3.5cm
 
|margin architecture or pubescence or shape=ciliate
 
|margin reflectance=nonlustrous
 
|margin width=thin
 
|node shape=swollen
 
|pair count=4;9
 
|petiole architecture=articulate
 
|petiole base count=abundant
 
|petiole base duration=persistent
 
|petiole coloration=dark purple;brown
 
|petiole fragility=brittle
 
|pinna length or size=longer than wide
 
|pinna shape=tapered;ovatelanceolate;deltate
 
|pinnule shape=lobed;crenate;entire
 
|projection coloration or reflectance=translucent
 
|projection count=lacking
 
|scale coloration=brown
 
|scale shape=linear-lanceolate;lanceolate
 
|segment shape=hairlike
 
|segment size or width=narrow
 
|sporangium life cycle=mature
 
|spore some measurement=39um;46um
 
|stem architecture or arrangement=compact
 
|stem orientation=erect;ascending
 
|vein tip size=enlarged
 
 
}}<!--
 
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-->[[Category:Treatment]][[Category:Woodsia]]
 
-->[[Category:Treatment]][[Category:Woodsia]]

Revision as of 13:17, 27 July 2019

Stems compact, erect to ascending, with abundant persistent petiole bases of ± equal length; scales uniformly brown, lanceolate. Leaves 4.5–25 × 1.2–3.5 cm. Petiole usually brown or dark purple when mature, articulate above base at swollen node, relatively brittle and easily shattered. Blade narrowly lanceolate, usually 2-pinnate proximally, lacking glands, never viscid; rachis usually with abundant hairs and scales. Pinnae ovate-lanceolate to deltate, longer than wide, abruptly tapered to a rounded or broadly acute apex; largest pinnae with 4–9 pairs of pinnules; abaxial surface with mixture of hairs and linear-lanceolate scales, adaxial surface with multicellular hairs concentrated along midrib. Pinnules entire or crenate, rarely shallowly lobed; margins nonlustrous, thin, ciliate with multicellular hairs, lacking translucent projections. Vein tips frequently enlarged to form whitish hydathodes visible adaxially. Indusia of narrow, hairlike segments, these uniseriate throughout, composed of cells many times longer than wide, usually surpassing mature sporangia. Spores averaging 39–46 µm. 2n = 82.


Phenology: Sporulating summer–early fall.
Habitat: Cliffs and rocky slopes, found on variety of substrates including serpentine
Elevation: 0–1500 m

Distribution

V2 349-distribution-map.gif

Greenland, Alta., B.C., Man., N.B., Nfld., N.W.T., N.S., Ont., Que., Sask., Yukon, Alaska, Conn., Ill., Iowa, Maine, Md., Mass., Mich., Minn., N.H., N.J., N.Y., N.C., Ohio, Pa., R.I., Vt., Va., W.Va., Wis., n Eurasia.

Discussion

Although generally separable by the characters given in the key, shade forms of Woodsia ilvensis with a reduced number of scales and hairs are occasionally misidentified as W. alpina. The morphologic distinctions between these species are further blurred by natural hybridization, which produces the intermediate triploid known as W. × gracilis. Some of the best characters for distinguishing these taxa are spore size and morphology. Spores average less than 46 µm in W. ilvensis, more than 46 µm in W. alpina, and are malformed and abortive in W. × gracilis. Woodsia ilvensis also hybridizes with W. oregana subsp. cathcartiana to form the sterile triploid W. × abbeae (F. S. Wagner 1987).

Selected References

None.

Lower Taxa

None.
... more about "Woodsia ilvensis"
Michael D. Windham +
(Linnaeus) R. Brown +
Acrostichum ilvense +
Rusty cliff fern +  and woodsie de l'île d'Elbe +
Greenland +, Alta. +, B.C. +, Man. +, N.B. +, Nfld. +, N.W.T. +, N.S. +, Ont. +, Que. +, Sask. +, Yukon +, Alaska +, Conn. +, Ill. +, Iowa +, Maine +, Md. +, Mass. +, Mich. +, Minn. +, N.H. +, N.J. +, N.Y. +, N.C. +, Ohio +, Pa. +, R.I. +, Vt. +, Va. +, W.Va. +, Wis. +  and n Eurasia. +
0–1500 m +
Cliffs and rocky slopes, found on variety of substrates including serpentine +
Sporulating summer–early fall. +
Trans. Linn. Soc. London, Bot. +
Woodsia ilvensis +
species +