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| − | --><span class="statement" id="st-d0_s0" data-properties="root reproduction"><b>Roots </b>not proliferous.</span> <span class="statement" id="st-d0_s1" data-properties="stem orientation;stem orientation;stem orientation;stem architecture"><b>Stems </b>ascending to erect, rarely branched;</span> <span class="statement" id="st-d0_s2" data-properties="scale coloration;scale coloration;scale coloration;scale shape;scale length;scale width;margin architecture or shape">scales dark-brown to blackish throughout, narrowly deltate, 2–4 × 0.25–0.45 mm, margins entire.</span> <span class="statement" id="st-d0_s3" data-properties="leaf variability;leaf reproduction;leaf height;leaf orientation;leaf reproduction"><b>Leaves </b>weakly subdimorphic, fertile leaves taller and more erect than sterile leaves.</span> <span class="statement" id="st-d0_s4" data-properties="petiole coloration;petiole coloration;petiole reflectance;petiole some measurement;petiole length"><b>Petiole </b>reddish or purplish brown throughout, lustrous, 1–10 cm, 1/5–1 times length of blade;</span> <span class="statement" id="st-d0_s5" data-properties="indument shape;scale coloration;scale coloration;scale coloration">indument of dark-brown to black scales, narrowly deltate at very base, grading into hairs distally.</span> <span class="statement" id="st-d0_s6" data-properties="blade variability;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade shape;blade length;blade atypical width;blade width;blade thickness;blade pubescence"><b>Blade </b>highly variable and typically irregularly shaped, narrowly deltate to lanceolate, pinnatifid or 1-pinnate in proximal 1/3, 2–20 × 1–6 (–13) cm, medium thick, sparsely pubescent adaxially only;</span> <span class="statement" id="st-d0_s7" data-properties="base architecture or shape">base ± truncate;</span> <span class="statement" id="st-d0_s8" data-properties="apex shape;apex shape;apex shape">apex acute to long-attenuate, apical buds borne occasionally but not known to root in nature.</span> <span class="statement" id="st-d0_s9" data-properties="rachis coloration;rachis coloration;rachis coloration;rachis coloration;rachis coloration;rachis coloration;rachis reflectance;rachis pubescence"><b>Rachis </b>reddish or purplish brown abaxially, fading to green distally, lustrous, glabrous.</span> <span class="statement" id="st-d0_s10" data-properties="pinna shape;pinna shape;pinna shape;pinna shape;pair count"><b>Pinnae </b>in 0–3 pairs, often irregular in size and shape, deltate to narrowly deltate;</span> <span class="statement" id="st-d0_s11" data-properties="proximal pinna atypical length;proximal pinna length;proximal pinna atypical width;proximal pinna width">proximal pinnae 5–30 (–80) × 3–10 (–15) mm;</span> <span class="statement" id="st-d0_s12" data-properties="base shape;base shape;base shape;base shape">base truncate to obtuse, auriculate on both sides;</span> <span class="statement" id="st-d0_s13" data-properties="margin architecture or shape;margin shape;margin shape">margins entire to finely serrate or crenulate;</span> <span class="statement" id="st-d0_s14" data-properties="apex shape;apex shape;apex shape;apex shape">apex obtuse to acute or occasionally attenuate.</span> <span class="statement" id="st-d0_s15" data-properties="vein prominence;vein fusion;vein architecture"><b>Veins </b>somewhat evident, mostly free, rarely anastomosing.</span> <span class="statement" id="st-d0_s16" data-properties="sorus atypical count;sorus count;lobe orientation;lobe orientation"><b>Sori </b>1–10 (–15+) pairs per pinna, on both acroscopic and basiscopic lobes.</span> <span class="statement" id="st-d0_s17" data-properties=""><b>Spores </b>malformed (sterile form) or 64 per sporangium (fertile form).</span> <span class="statement" id="st-d0_s18" data-properties="spore architecture;spore count;2n chromosome count;2n chromosome count">2n = 72, 144.</span><!-- | + | --><span class="statement" id="st-undefined" data-properties=""><b>Roots </b>not proliferous. <b>Stems</b> ascending to erect, rarely branched; scales dark brown to blackish throughout, narrowly deltate, 2–4 × 0.25–0.45 mm, margins entire. <b>Leaves</b> weakly subdimorphic, fertile leaves taller and more erect than sterile leaves. <b>Petiole</b> reddish or purplish brown throughout, lustrous, 1–10 cm, 1/5–1 times length of blade; indument of dark brown to black scales, narrowly deltate at very base, grading into hairs distally. <b>Blade</b> highly variable and typically irregularly shaped, narrowly deltate to lanceolate, pinnatifid or 1-pinnate in proximal 1/3, 2–20 × 1–6(–13) cm, medium thick, sparsely pubescent adaxially only; base ± truncate; apex acute to long-attenuate, apical buds borne occasionally but not known to root in nature. <b>Rachis</b> reddish or purplish brown abaxially, fading to green distally, lustrous, glabrous. <b>Pinnae</b> in 0–3 pairs, often irregular in size and shape, deltate to narrowly deltate; proximal pinnae 5–30(–80) × 3–10(–15) mm; base truncate to obtuse, auriculate on both sides; margins entire to finely serrate or crenulate; apex obtuse to acute or occasionally attenuate. <b>Veins</b> somewhat evident, mostly free, rarely anastomosing. <b>Sori</b> 1–10(–15+) pairs per pinna, on both acroscopic and basiscopic lobes. <b>Spores</b> malformed (sterile form) or 64 per sporangium (fertile form). <b>2n</b> = 72, 144.</span><!-- |
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| | |publication year=1865 | | |publication year=1865 |
| | |special status= | | |special status= |
| − | |source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_341.xml | + | |source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_341.xml |
| | |genus=Asplenium | | |genus=Asplenium |
| | |species=Asplenium ebenoides | | |species=Asplenium ebenoides |
| − | |2n chromosome count=144;72
| |
| − | |apex shape=obtuse;acute or occasionally attenuate
| |
| − | |base architecture or shape=truncate
| |
| − | |base shape=auriculate;truncate;obtuse
| |
| − | |blade atypical width=6cm;13cm
| |
| − | |blade length=2cm;20cm
| |
| − | |blade pubescence=pubescent
| |
| − | |blade shape=narrowly deltate;lanceolate pinnatifid or 1-pinnate
| |
| − | |blade thickness=medium
| |
| − | |blade variability=variable
| |
| − | |blade width=1cm;6cm
| |
| − | |indument shape=deltate
| |
| − | |leaf height=taller
| |
| − | |leaf orientation=erect
| |
| − | |leaf reproduction=sterile;fertile
| |
| − | |leaf variability=subdimorphic
| |
| − | |lobe orientation=basiscopic;acroscopic
| |
| − | |margin architecture or shape=entire;entire
| |
| − | |margin shape=crenulate;serrate
| |
| − | |pair count=0;3
| |
| − | |petiole coloration=purplish brown;reddish
| |
| − | |petiole length=1/5-1 times length of blade
| |
| − | |petiole reflectance=lustrous
| |
| − | |petiole some measurement=1cm;10cm
| |
| − | |pinna shape=deltate;narrowly deltate
| |
| − | |proximal pinna atypical length=30mm;80mm
| |
| − | |proximal pinna atypical width=10mm;15mm
| |
| − | |proximal pinna length=5mm;30mm
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| − | |proximal pinna width=3mm;10mm
| |
| − | |rachis coloration=fading;green
| |
| − | |rachis pubescence=glabrous
| |
| − | |rachis reflectance=lustrous
| |
| − | |root reproduction=not proliferous
| |
| − | |scale coloration=dark-brown;black
| |
| − | |scale length=2mm;4mm
| |
| − | |scale shape=deltate
| |
| − | |scale width=0.25mm;0.45mm
| |
| − | |sorus atypical count=10;15
| |
| − | |sorus count=1;10
| |
| − | |spore architecture=malformed
| |
| − | |spore count=64
| |
| − | |stem architecture=branched
| |
| − | |stem orientation=ascending;erect
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| − | |vein architecture=anastomosing
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| − | |vein fusion=free
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| − | |vein prominence=evident
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| | -->[[Category:Treatment]][[Category:Asplenium]] | | -->[[Category:Treatment]][[Category:Asplenium]] |
Roots not proliferous. Stems ascending to erect, rarely branched; scales dark brown to blackish throughout, narrowly deltate, 2–4 × 0.25–0.45 mm, margins entire. Leaves weakly subdimorphic, fertile leaves taller and more erect than sterile leaves. Petiole reddish or purplish brown throughout, lustrous, 1–10 cm, 1/5–1 times length of blade; indument of dark brown to black scales, narrowly deltate at very base, grading into hairs distally. Blade highly variable and typically irregularly shaped, narrowly deltate to lanceolate, pinnatifid or 1-pinnate in proximal 1/3, 2–20 × 1–6(–13) cm, medium thick, sparsely pubescent adaxially only; base ± truncate; apex acute to long-attenuate, apical buds borne occasionally but not known to root in nature. Rachis reddish or purplish brown abaxially, fading to green distally, lustrous, glabrous. Pinnae in 0–3 pairs, often irregular in size and shape, deltate to narrowly deltate; proximal pinnae 5–30(–80) × 3–10(–15) mm; base truncate to obtuse, auriculate on both sides; margins entire to finely serrate or crenulate; apex obtuse to acute or occasionally attenuate. Veins somewhat evident, mostly free, rarely anastomosing. Sori 1–10(–15+) pairs per pinna, on both acroscopic and basiscopic lobes. Spores malformed (sterile form) or 64 per sporangium (fertile form). 2n = 72, 144.
Habitat: Conglomerate boulders
Elevation: 70 m
Discussion
The above description applies to the sterile hybrid Asplenium platyneuron × rhizophyllum and its allopolyploid derivative. The allotetraploid form is known only from Hale County, Alabama, where it occurs with A. platyneuron (but not with A. rhizophyllum) on conglomerate boulders (K. S. Walter et al. 1982). The sterile diploid form of A. ebenoides occurs at elevations of 70 to 500 m within the region where the ranges of the parental species overlap, always occurring with both parents on limestone, sandstone, or other rock strata. A hybrid between the allopolyploid and A. platyneuron [A. × boydstoniae (K. S. Walter) J. W. Short] was discovered at Havana Glen. An unnamed hybrid between the sterile diploid (presumably via unreduced spores) and A. rhizophyllum is known from West Virginia and Missouri (K. S. Walter et al. 1982).
This fern has been pivotal in the study of fern hybridization. Called the "most famous hybrid fern," it was one of the first crosses to be synthesized deliberately in culture (M. Slosson 1902) and the first to be converted from the sterile diploid state to the fertile tetraploid state experimentally (W. H. Wagner Jr. and R. S. Whitmire 1957).
Selected References
None.
