Hamamelidaceae

Trees or shrubs, deciduous; pubescence stellate or sometimes simple. Dormant buds naked or scaly. Leaves alternate, simple, petiolate; stipules early deciduous. Leaf-blade unlobed or deeply (3-) 5-7-lobed, pinnately or palmately veined. Inflorescences axillary or terminal, capitate or spicate to ± racemose, bracteate. Flowers bisexual or unisexual; perianth epigynous, 1-2-seriate, 4-5-merous, often reduced or absent, hypanthium present; calyx persistent, 4-5 (-7) -lobed, or absent; petals distinct or absent; stamens 4-34; anthers basifixed, latrorse, 2-locular, longitudinally dehiscent by simple slit or by 1-2 vertical valves; pistil 1, 2-carpellate; ovary half-inferior, 2-locular; placentation axile; ovules 1 (-2) in each locule and apical, or many, anatropous, pendent; styles and stigmas persistent, 2 each, erect or contorted and recurved. Fruits capsular, with leathery exocarp and bony endocarp, loculicidal and 2-seeded or septicidal with most of many seeds per locule aborted; 1-2 viable seeds per locule (Liquidambar). Seeds fusiform, bony, lustrous, hilum apical, light colored, or seeds winged, hilum lateral (Liquidambar).

Distribution

Temperate to tropical regions, e North America, Mexico, Central America, e Asia, Africa (including Madagascar), Pacific Islands, and Australia

Discussion

Genera ca. 31, species ca. 100 (3 genera, 5 species in the flora).

Selected References

Bogle, A. L. 1970. Floral morphology and vascular anatomy of the Hamamelidaceae: The apetalous genera of Hamamelidoideae. J. Arnold Arbor. 51: 310-366.

Britton, N. L. 1905. Hamamelidaceae. In: N. L. Britton et al., eds. 1905+. North American Flora.... 47+ vols. New York. Vol. 22, pp. 185-187.

Endress, P. K. 1987. Aspects of evolutionary differentiation of the Hamamelidaceae and the lower Hamamelidae. Pl. Syst. Evol. 162: 193-211.

Endress, P. K. 1989. A suprageneric taxonomic classification of the Hamamelidaceae. Taxon 38(3): 371-376.

Endress, P. K. 1989b. Phylogenetic relationships in the Hamamelidoideae. In: P. R. Crane and S. Blackmore, eds. 1989. Evolution, Systematics, and Fossil History of the Hamamelidae. 2 vols. Oxford. Vol. 1, pp. 227-248. [Syst. Assoc. Special Vol. 40A, B.]

Ernst, W. R. 1963. The genera of Hamamelidaceae and Platanaceae in the southeastern United States. J. Arnold Arbor. 44: 193-210.

Godfrey, R. K. 1988. Trees, Shrubs, and Woody Vines of Northern Florida and Adjacent Georgia and Alabama. Athens, Ga.

Goldblatt, P. and P. K. Endress. 1977. Cytology and evolution in Hamamelidaceae. J. Arnold Arbor. 58: 67-71.

Lower Taxa

Fothergilla, Hamamelis, Liquidambar

Illustrations

	Family ⠉	Taxon	Illustrator ⠉
	Hamamelidaceae Hamamelidaceae Hamamelidaceae Hamamelidaceae	Liquidambar styraciflua Fothergilla gardenii Hamamelis virginiana Fothergilla major	John Myers John Myers John Myers John Myers

Key

1	Trees with simple hairs; leaf blade palmately (3-)5(-7)-lobed and - veined; flowers unisexual, terminal, the staminate in ± racemose heads, each head a clustered mass of numerous stamens, pistillate inflorescences many flowered; capsules septicidal, fused at base into long-pedunculate, globose, echinate heads; viable seeds winged, most seeds aborted.	Liquidambar
1	Shrubs or small trees with stellate pubescence; leaf blade unlobed, pinnately veined; flowers bisexual, in axillary, few-flowered clusters or in many-flowered, elongate spikes; capsules loculicidal, not fused into heads; viable seeds not winged.	> 2
2	Inflorescences axillary, (1-)3(-5)-flowered clusters; calyx lobes 4, reflexed; petals 4, liguliform, yellow or reddish, deep red to orange; stamens 4, very short; staminodes 4; capsules solitary or 2-3 together.	Hamamelis
2	Inflorescences terminal, elongate, many-flowered spikes; calyx lobes 5-7(-9), erect; petals absent; stamens 12-34; filaments white, 4-17 mm; staminodes absent; capsules in groups of more than 3.	Fothergilla