View source for Athyrium

{{Treatment/ID
|accepted_name=Athyrium
|accepted_authority=Roth
|publications={{Treatment/Publication
|title=Tent. Fl. Germ.
|place=3(1,1): 31, 58. 1799
|year=1799
}}
|common_names=Lady fern
|basionyms=
|synonyms=
|hierarchy=Dryopteridaceae;Athyrium
|hierarchy_nav=<div class="higher-taxa"><div class="higher-taxon"><small>family</small>[[Dryopteridaceae]]</div><div class="higher-taxon"><small>genus</small>[[Athyrium]]</div></div>
|etymology=Greek athyros, doorless; the sporangia only tardily push back the outer edge of the indusium
|volume=Volume 2
|mention_page=
|treatment_page=
}}<!--

--><span class="statement" id="st-d0_s0" data-properties="plant growth form or habitat"><b>Plants </b>generally terrestrial.</span> <span class="statement" id="st-d0_s1" data-properties="stem orientation;stem orientation;stolon count"><b>Stems </b>short-creeping or ascending, stolons absent.</span> <span class="statement" id="st-d0_s2" data-properties="leaf architecture;leaf condition"><b>Leaves </b>monomorphic, usually dying back in winter.</span> <span class="statement" id="st-d0_s3" data-properties="petiole length;petiole length;base shape;base architecture or shape;base duration"><b>Petiole </b>± 0.5 times length of blade or less, base swollen and dentate, persisting as trophopod over winter or not;</span> <span class="statement" id="st-d0_s4" data-properties="vascular-bundle count;vascular-bundle position;vascular-bundle shape">vascular-bundles 2, lateral, lunate in cross-section.</span> <span class="statement" id="st-d0_s5" data-properties="blade shape;blade shape;blade shape;blade shape;blade shape;blade size;blade growth form or texture;apex arrangement;apex shape"><b>Blade </b>lanceolate to elliptic or oblanceolate, 1–3-pinnate-pinnatifid, gradually reduced distally to confluent, pinnatifid apex, herbaceous.</span> <span class="statement" id="st-d0_s6" data-properties="pinna architecture;segment margin shape;segment margin shape"><b>Pinnae </b>not articulate to rachis, segment margins serrulate or crenate;</span> <span class="statement" id="st-d0_s7" data-properties="proximal pinna size;proximal pinna architecture;proximal pinna architecture;proximal pinna architecture;proximal pinna architecture or shape">proximal pinnae often reduced, sessile to short-petiolulate, ± equilateral;</span> <span class="statement" id="st-d0_s8" data-properties="costa architecture;groove architecture">costae adaxially grooved, grooves continuous from rachis to costae to costules;</span> <span class="statement" id="st-d0_s9" data-properties="indument count;scale shape;scale shape;scale shape;gland architecture">indument absent or of linear to lanceolate scales or 1-celled glands abaxially.</span> <span class="statement" id="st-d0_s10" data-properties="vein fusion;vein architecture;vein shape"><b>Veins </b>free, simple or forked.</span> <span class="statement" id="st-d0_s11" data-properties="between midrib and margin row count"><b>Sori </b>in 1 row between midrib and margin, round to elongate, straight or hooked at distal end, or horseshoe-shaped;</span> <span class="statement" id="st-d0_s12" data-properties="sorus shape;sorus shape;sorus shape;sorus course;sorus shape;sorus shape;sorus shape;sorus duration;sorus fixation;sorus fixation;sinus size or width;indusium count">indusia shaped like sori, persistent, attached laterally or with narrow sinus, or indusia absent.</span> <span class="statement" id="st-d0_s13" data-properties=""><b>Spores </b>brownish, rugose.</span> <span class="statement" id="st-d0_s14" data-properties="spore coloration;spore relief;x chromosome count">x = 40.</span><!--

-->{{Treatment/Body
|distribution=Worldwide.
|discussion=<p>In species outside the flora stems are sometimes long-creeping to erect, with leaves radially or dorsiventrally arranged.</p><!--
--><p>Species about 180 (2 in the flora).</p>
|tables=
|references={{Treatment/Reference
|id=johnson1986c
|text=Johnson, D. M. 1986b. Trophopods in North American species of Athyrium (Aspleniaceae). Syst. Bot. 11: 26--31.
}}{{Treatment/Reference
|id=kato1977a
|text=Kato, M. 1977. Classification of Athyrium and allied genera of Japan. Bot. Mag. (Tokyo) 90: 23--40.
}}{{Treatment/Reference
|id=liew1972a
|text=Liew, F. S. 1972. Numerical taxonomic studies on North American lady ferns and their allies. Taiwania 17: 190--221.
}}
}}<!--

--><div class="treatment-key">
==Key==
<div class="treatment-key-group">
  
{| class="wikitable fna-keytable"
|-id=key-0-1
|1
|Sori round, submarginal; indusia much reduced or usually absent.
|[[Athyrium alpestre var. americanum|Athyrium alpestre var. americanum]]
|-id=key-0-1
|1
|Sori elongate or hooked, medial; indusia well developed.
|[[Athyrium filix-femina|Athyrium filix-femina]]
|}
</div></div><!--

-->{{#Taxon:
name=Athyrium
|author=Masahiro Kato
|authority=Roth
|rank=genus
|parent rank=family
|synonyms=
|basionyms=
|family=Dryopteridaceae
|distribution=Worldwide.
|reference=johnson1986c;kato1977a;liew1972a
|publication title=Tent. Fl. Germ.
|publication year=1799
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_540.xml
|genus=Athyrium
|apex arrangement=confluent
|apex shape=pinnatifid
|base architecture or shape=dentate
|base duration=persisting
|base shape=swollen
|between midrib and margin row count=1
|blade growth form or texture=herbaceous
|blade shape=1-3-pinnate-pinnatifid;lanceolate;elliptic or oblanceolate
|blade size=reduced
|costa architecture=grooved
|gland architecture=1-celled
|groove architecture=continuous
|indument count=absent
|indusium count=absent
|leaf architecture=monomorphic
|leaf condition=dying
|petiole length=0.5 times length of blade;0.5 times length of blade
|pinna architecture=not articulate
|plant growth form or habitat=terrestrial
|proximal pinna architecture=sessile;short-petiolulate
|proximal pinna architecture or shape=equilateral
|proximal pinna size=reduced
|scale shape=linear;lanceolate
|segment margin shape=crenate;serrulate
|sinus size or width=narrow
|sorus course=straight
|sorus duration=persistent
|sorus fixation=with narrow sinus;attached
|sorus shape=indusia-shaped;horseshoe-shaped;hooked;round;elongate
|spore coloration=brownish
|spore relief=rugose
|stem orientation=ascending;short-creeping
|stolon count=absent
|vascular-bundle count=2
|vascular-bundle position=lateral
|vascular-bundle shape=lunate
|vein architecture=simple
|vein fusion=free
|vein shape=forked
|x chromosome count=40
}}<!--

-->[[Category:Treatment]][[Category:Dryopteridaceae]]