Pseudoleskea radicosa

(Mitten) Macoun & Kindberg

Cat. Canad. Pl., Musci, 181. 1892.

Basionym: Hypnum radicosum Mitten J. Linn. Soc., Bot. 8: 31. 1864
Synonyms: Lescuraea radicosa (Mitten) Mönke meyer
Treatment appears in FNA Volume 28. Treatment on page 359. Mentioned on page 356, 357.

Plants medium-sized to large, in thick mats, green, yellow-green, or orange-green. Stems with branches robust, julaceous, apices usually curving up; central strand present; paraphyllia many, filamentous to foliose, branched. Leaves appressed to somewhat erect when dry, erect-spreading when moist, glossy or dull, ovate to ovate-lanceolate, asymmetric, usually falcate to falcate-secund, 0.6–2(–2.4) mm; margins narrowly recurved to mid leaf or to near acumen; apex abruptly acute to short- or long-acuminate, hair-point absent; costa subpercurrent to percurrent, yellow-green, sometimes sinuate; alar cells transversely elongate to quadrate, region small to medium-sized; medial laminal cells homogeneous, short-rhomboidal, elliptic, or fusiform, to 40 µm, 2–3(–4):1, pellucid to opaque, prorate to near base, lumina larger than 10 µm, walls thin or rarely firm, not pitted; apical cells 2–3:1; juxtacostal cells somewhat shorter than more distal cells, walls not pitted. Capsule erect to suberect, symmetric, 1–2 mm; endostome basal membrane 1/4–1/3 exostome length, segments shorter than exostome, cilia usually present, 1 or 2, sometimes reduced. Spores 16–22 µm.

Distribution

V28 561-distribution-map.gif

North America, Eurasia, Atlantic Islands (Iceland).

Discussion

Varieties 3 (3 in the flora).

Pseudoleskea radicosa is a common and variable species found at moderate to high elevations; the capsules mature in summer. Two of the three varieties are quite distinctive. Pseudoleskea radicosa is closely related to P. incurvata but differs in the longer, more homogeneous, thin-walled, and wider laminal cells. Combined ecological, molecular and morphological studies of the varieties are needed to determine their distinctiveness.

Selected References

None.

Key

1 Plants large; stem apices hooked; branches few or absent; leaves 1.6-2.3 mm, strongly falcate-secund, pluriplicate; apices long-acuminate. Pseudoleskea radicosa var. denudata
1 Plants small to medium-sized; stem apices not distinctly hooked; branches many; leaves usually less than 1.6 mm, straight, falcate, or falcate-secund, 2-plicate; apices predominantly short-acuminate to acute > 2
2 Plants medium-sized; leaves weakly to moderately concave, not incurved, 1.2-1.7 mm; apices usually long-acuminate; medial laminal cells opaque. Pseudoleskea radicosa var. radicosa
2 Plants small; leaves strongly concave, often incurved, 0.8-1.3 mm; apices acute to short-acuminate; medial laminal cells pellucid. Pseudoleskea radicosa var. compacta