Pellaea glabella subsp. occidentalis

(E. E. Nelson) Windham

Contr. Univ. Michigan Herb. 19: 39. 1993.

Endemic
Basionym: Pellaea atropurpurea var. occidentalis E. E. Nelson Fern Bull. 7: 30. 1899
Synonyms: Pellaea glabella var. nana (Richardson) Cody Pellaea glabella var. occidentalis (E. E. Nelson) Butters Pellaea occidentalis Pellaea pumila
Treatment appears in FNA Volume 2.

Ultimate segments glabrous; sporangia containing 64 spores; spores averaging 38–52 µm diam. 2n = 58.


Phenology: Sporulating summer–fall.
Habitat: Calcareous cliffs and ledges, usually on limestone
Elevation: 500–2800 m

Distribution

V2 625-distribution-map.gif

Man., Sask., Mont., N.Dak., S.Dak., Utah, Wyo.

Discussion

Most recent treatments refer to this taxon as Pellaea glabella var. occidentalis. It appears to be synonymous, however, with P. glabella var. nana, which was proposed 76 years earlier and has priority if the taxon is treated at varietal rank. This diploid (treated here as a subspecies) often shows prominent articulation lines near the base of the petiole, and plants with less divided leaves are occasionally misidentified as P. breweri. At least some of the sporangia of subsp. occidentalis are long-stalked, however, whereas those of P. breweri are sessile or subsessile.

Selected References

None.

Lower Taxa

None.
Michael D. Windham +
(E. E. Nelson) Windham +
Pellaea atropurpurea var. occidentalis +
Man. +, Sask. +, Mont. +, N.Dak. +, S.Dak. +, Utah +  and Wyo. +
500–2800 m +
Calcareous cliffs and ledges, usually on limestone +
Sporulating summer–fall. +
Contr. Univ. Michigan Herb. +
Pellaea glabella var. nana +, Pellaea glabella var. occidentalis +, Pellaea occidentalis +  and Pellaea pumila +
Pellaea glabella subsp. occidentalis +
Pellaea glabella +
subspecies +