Ephemerum

Stems absent. Leaves scarcely contorted when dry; costa well-developed, except generally absent or weak in E. serratum; laminal cells lax in the proximal half of the leaf, in the distal half firmer, smaller, and usually papillose. Sexual condition dioicous or autoicous, commonly rhizautoicous. Capsule cleistocarpous, ovoid and distinctly apiculate, more than 0.5 mm from base to apex; exothecium of 2 cell layers; columella present in maturing capsules of E. cohaerens, but in other species visible only as pigmented areas at base and tip of the spore sac. Calyptra fugacious, cucullate, sometimes lobed or torn at the base, covering the distal 2/3 of the capsule, mostly more than 0.2 mm.

Distribution

Nearly worldwide in temperate and tropical regions.

Discussion

Species ca. 28 (4 in the flora).

All Ephemerum taxa in the flora have the same chromosome number, n = 27. Mixed colonies are common, although hybrid forms are not evident. To a large extent, the species occupy similar ranges.

Selected References

None.

Lower Taxa

Ephemerum cohaerens, Ephemerum crassinervium, Ephemerum serratum, Ephemerum spinulosum

Key

1	Leaves ecostate or in the distal two-thirds very weakly costate.	Ephemerum serratum
1	Leaves distinctly costate, at least distal to the middle	> 2
2	Distal laminal cells spinulose or spinose.	Ephemerum spinulosum
2	Distal laminal cells smooth or papillose	> 3
3	Distal laminal cells smooth, in ± diagonal rows.	Ephemerum cohaerens
3	Distal laminal cells papillose in ± vertical rows.	Ephemerum crassinervium