Ptychostomum turbinatum

Plants in dense or open turfs, dull green, yellow-green, or brownish. Stems (1–)2–4(–6) cm, fertile stems comose, innovations evenly foliate. Leaves dull green to yellow-green, crowded, strongly contorted to shrunken when dry, often broadly ovate-lanceolate, somewhat concave, 1–3 mm, gradually enlarged toward stem apex; base green, not or weakly decurrent; margins plane or occasionally weakly revolute proximally, limbidium strong, in 2 or 3 rows; apex acute to acuminate; costa percurrent to short-excurrent, awn slender; proximal laminal cells rectangular, 3–4:1; medial and distal cells 18–25(–30) µm wide, 2–3:1, walls thin. Specialized asexual reproduction absent. Sexual condition dioicous. Seta brown or red-brown, 1–3(–4) cm, slender, straight to somewhat flexuose. Capsule pale yellow to yellow-brown, becoming black with age, turbinate to short-pyriform, symmetric, 3–5 mm, mouth yellow; operculum conic, apiculate; peristome well developed; exostome teeth yellow to orange basally, hyaline distally, lamellae straight, pores absent near base along mid line; endostome not adherent to exostome, basal membrane 1/2 exostome height, segments with ovate perforations, cilia appendiculate to nodose. Spores 18–22 µm, yellow to brown.

Phenology: Capsules mature Jun–Aug.
Habitat: Wet soil in calcareous wetlands
Elevation: low to high elevations (0-3500 m)

Distribution

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Greenland, Alta., B.C., Nfld. and Labr., N.W.T., Yukon, Alaska, Calif., Colo., Idaho, Mont., Nev., Oreg., Utah, Wash., Wyo., South America (Argentina, Chile, Ecuador, Peru), Eurasia (including Turkey), Africa.

Discussion

Ptychostomum turbinatum is a circumpolar arctic-boreal to temperate species. The dull green to yellow-green shoots, strongly shrunken ovate-lanceolate leaves, and turbinate capsule are diagnostic. Old capsules often turn black and become strongly constricted just below the mouth. The stems are sparsely radiculose, sometimes with clusters of rhizoids arising from the proximal leaf axils. Ptychostomum schleicheri is much larger, pale yellow distally, with wider leaves and much wider distal laminal cells. Large specimens, 8–12 cm, with long narrow ovate-lanceolate leaves and elongate-pyriform capsules, represent an undescribed species.

Selected References

None.

Lower Taxa

None.