Difference between revisions of "Pellaea ternifolia"

(Cavanilles) Link

Fil. Spec. 59. 1841.

Basionym: Pteris ternifolia Cavanilles
Treatment appears in FNA Volume 2. Treatment on page 180.
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--><span class="statement" id="st-d0_s0" data-properties="stem architecture or arrangement;stem orientation;stem fragility or size;stem diameter"><b>Stems </b>compact, ascending, stout, 5–10 mm diam.;</span> <span class="statement" id="st-d0_s1" data-properties="scale coloration;scale shape;scale width;center coloration;center width;margin coloration;margin width;margin architecture or shape">scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate.</span> <span class="statement" id="st-d0_s2" data-properties="leaf architecture;leaf arrangement or growth form;leaf some measurement"><b>Leaves </b>monomorphic, clustered on stem, 10–50 cm;</span> <span class="statement" id="st-d0_s3" data-properties="crozier pubescence">croziers sparsely to densely villous.</span> <span class="statement" id="st-d0_s4" data-properties="petiole coloration;petiole coloration;petiole reflectance;petiole shape;petiole shape;articulation line prominence"><b>Petiole </b>black or dark purple, lustrous, rounded or slightly flattened adaxially, without prominent articulation lines.</span> <span class="statement" id="st-d0_s5" data-properties="blade shape;blade shape;blade shape;blade shape;blade width"><b>Blade </b>linear to ovate, deeply pinnate-pinnatifid proximally, 2.5–8 cm wide;</span> <span class="statement" id="st-d0_s6" data-properties="rachis coloration;rachis coloration;rachis course;rachis shape;rachis pubescence;rachis pubescence">rachis black or purple throughout, straight, often flattened adaxially, glabrous or villous.</span> <span class="statement" id="st-d0_s7" data-properties="pinna orientation;pinna architecture;rachis orientation;rachis shape"><b>Pinnae </b>perpendicular to rachis or slightly ascending, not decurrent on rachis, ternate at base of leaf;</span> <span class="statement" id="st-d0_s8" data-properties="costa count">costae absent.</span> <span class="statement" id="st-d0_s9" data-properties="ultimate segment shape;ultimate segment some measurement;ultimate segment texture;ultimate segment pubescence;ultimate segment pubescence;ultimate segment pubescence"><b>Ultimate </b>segments linear-oblong, 10–40 mm, leathery, glabrous to sparsely villous abaxially on midrib;</span> <span class="statement" id="st-d0_s10" data-properties="margin orientation;segment reproduction;abaxial surface count;abaxial surface count;border coloration;border architecture or shape">margins recurved on fertile segments, rarely covering more than 1/2 abaxial surface, borders whitish, entire;</span> <span class="statement" id="st-d0_s11" data-properties="apex shape">apex mucronate.</span> <span class="statement" id="st-d0_s12" data-properties="vein prominence"><b>Veins </b>of ultimate segments obscure.</span> <span class="statement" id="st-d0_s13" data-properties="sporangium architecture;spore count"><b>Sporangia </b>long-stalked, containing 64 spores, not intermixed with farina-producing glands.</span><!--
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--><span class="statement" id="st-undefined" data-properties=""><b>Stems </b>compact, ascending, stout, 5–10 mm diam.; scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. <b>Leaves</b> monomorphic, clustered on stem, 10–50 cm; croziers sparsely to densely villous. <b>Petiole</b> black or dark purple, lustrous, rounded or slightly flattened adaxially, without prominent articulation lines. <b>Blade</b> linear to ovate, deeply pinnate-pinnatifid proximally, 2.5–8 cm wide; rachis black or purple throughout, straight, often flattened adaxially, glabrous or villous. <b>Pinnae</b> perpendicular to rachis or slightly ascending, not decurrent on rachis, ternate at base of leaf; costae absent. <b>Ultimate</b> segments linear-oblong, 10–40 mm, leathery, glabrous to sparsely villous abaxially on midrib; margins recurved on fertile segments, rarely covering more than 1/2 abaxial surface, borders whitish, entire; apex mucronate. <b>Veins</b> of ultimate segments obscure. <b>Sporangia</b> long-stalked, containing 64 spores, not intermixed with farina-producing glands.</span><!--
  
 
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|publication year=1841
 
|publication year=1841
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_170.xml
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|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-data-curation.git/src/9216fc802291cd3df363fd52122300479582ede7/coarse_grained_fna_xml/V2/V2_170.xml
 
|genus=Pellaea
 
|genus=Pellaea
 
|species=Pellaea ternifolia
 
|species=Pellaea ternifolia
|abaxial surface count=/2;1
 
|apex shape=mucronate
 
|articulation line prominence=prominent
 
|blade shape=pinnate-pinnatifid;linear;ovate
 
|blade width=2.5cm;8cm
 
|border architecture or shape=entire
 
|border coloration=whitish
 
|center coloration=black
 
|center width=thick
 
|costa count=absent
 
|crozier pubescence=villous
 
|leaf architecture=monomorphic
 
|leaf arrangement or growth form=clustered
 
|leaf some measurement=10cm;50cm
 
|margin architecture or shape=erose-dentate
 
|margin coloration=brown
 
|margin orientation=recurved
 
|margin width=thin
 
|petiole coloration=dark purple;black
 
|petiole reflectance=lustrous
 
|petiole shape=flattened;rounded
 
|pinna architecture=ternate
 
|pinna orientation=perpendicular
 
|rachis coloration=purple;black
 
|rachis course=straight
 
|rachis orientation=ascending
 
|rachis pubescence=villous;glabrous
 
|rachis shape=not decurrent;flattened
 
|scale coloration=bicolored
 
|scale shape=linear-subulate
 
|scale width=0.1mm;0.3mm
 
|segment reproduction=fertile
 
|sporangium architecture=long-stalked
 
|spore count=64
 
|stem architecture or arrangement=compact
 
|stem diameter=5mm;10mm
 
|stem fragility or size=stout
 
|stem orientation=ascending
 
|ultimate segment pubescence=glabrous;sparsely villous
 
|ultimate segment shape=linear-oblong
 
|ultimate segment some measurement=10mm;40mm
 
|ultimate segment texture=leathery
 
|vein prominence=obscure
 
 
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Revision as of 13:12, 27 July 2019

Stems compact, ascending, stout, 5–10 mm diam.; scales bicolored, linear-subulate, 0.1–0.3 mm wide, centers black, thick, margins brown, thin, erose-dentate. Leaves monomorphic, clustered on stem, 10–50 cm; croziers sparsely to densely villous. Petiole black or dark purple, lustrous, rounded or slightly flattened adaxially, without prominent articulation lines. Blade linear to ovate, deeply pinnate-pinnatifid proximally, 2.5–8 cm wide; rachis black or purple throughout, straight, often flattened adaxially, glabrous or villous. Pinnae perpendicular to rachis or slightly ascending, not decurrent on rachis, ternate at base of leaf; costae absent. Ultimate segments linear-oblong, 10–40 mm, leathery, glabrous to sparsely villous abaxially on midrib; margins recurved on fertile segments, rarely covering more than 1/2 abaxial surface, borders whitish, entire; apex mucronate. Veins of ultimate segments obscure. Sporangia long-stalked, containing 64 spores, not intermixed with farina-producing glands.

Distribution

North America, Mexico, Central America, South America, Pacific Islands in Hawaii.

Discussion

Subspecies 3 (3 in the flora).

Pellaea ternifolia is represented in the flora by three morphologically and chromosomally distinct taxa. These discrete genetic entities also show a tendency toward geographic isolation and are treated here as subspecies. Diploid populations referred to P. ternifolia subsp. ternifolia are scattered from Texas through Mexico to South America. The pubescent tetraploid (P. ternifolia subsp. villosa) follows the Sierra Madre Oriental from Puebla, Mexico, north to Texas; the glabrous tetraploid (P. ternifolia subsp. arizonica) occurs in Arizona, Texas, and northern Mexico. Isozyme and chromosome studies suggest that both tetraploids are segmental allopolyploids produced by hybridization between subsp. ternifolia and other (as yet unidentified) diploid elements within P. ternifolia.

Selected References

None.

Key

1 Rachises villous, especially in axils of pinnae; pinnae with hairs scattered along main veins abaxially. Pellaea ternifolia subsp. villosa
1 Rachises glabrous or with a few widely scattered hairs; pinnae completely glabrous. > 2
2 Largest ultimate segments (excluding terminal pinnae) usually less than 18 mm; distal portion of petioles grooved or flattened adaxially; spores usually 39–45 µm diam. Pellaea ternifolia subsp. ternifolia
2 Largest ultimate segments (excluding terminal pinnae) usually more than 18 mm; distal portion of petioles rounded or slightly flattened adaxially; spores usually 46–53 µm diam. Pellaea ternifolia subsp. arizonica