Tortula acaulon

Leaves ovate to oblong-lanceolate, apex acute, mucronate to short-awned, occasionally comparatively long-awned, margins recurved proximally to nearly plane, not or weakly bordered distally with slightly thicker-walled cells; costa excurrent, lacking an adaxial pad of cells, distally narrow, 2(–3) cells across adaxial surface; distal laminal cells distally quadrate-hexagonal, width 13–17 µm, 1:1, weakly simply papillose or smooth. Sexual condition autoicous. Sporophytes immersed. Seta very short. Capsule cleistocarpic, spheric to very short-ellipsoid, erect, mostly 0.9–1.3 mm; peristome absent; operculum not differentiated. Spores (25–)33–40 µm, spheric, densely papillose.

Phenology: Capsules mature winter–spring.
Habitat: Soil, lawns, fields, banks
Elevation: low to moderate elevations

Distribution

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Alta., B.C., N.S., Ont., Que., Sask., Alaska, Ariz., Ark., Calif., Colo., Conn., Del., Fla., Idaho, Ill., Ind., Iowa, Kans., La., Md., Mich., Minn., Mo., Mont., Nebr., Nev., N.J., N.Mex., N.Y., N.C., Ohio, Okla., Oreg., Pa., S.Dak., Tenn., Tex., Utah., Va., Wash., W.Va., Wis., Mexico, Europe, Asia, n Africa, Pacific Islands (New Zealand).

Discussion

The varieties pilifera and schreberiana are not recognized here. The former name may be used for plants with long awns, and the latter for robust specimens ranging to 1 cm tall, but intergrades appear to be common.

Selected References

None.

Lower Taxa

None.