Scolymus

Annuals, biennials, or perennials, 10–80(–200+) cm; taprooted. Stems usually 1, erect, branched distally, glabrous or hairy (internodes winged, margins spiny, faces ± scabrellous and/or sparsely arachnose). Leaves basal and cauline (mostly cauline at flowering); basal ± petiolate, distal sessile; blades oblong or lanceolate to oblanceolate or linear, margins pinnately lobed to dentate, usually strongly spiny (faces ± scabrellous and/or sparsely arachnose). Heads borne singly (axillary and terminal) or in ± spiciform arrays. Peduncles usually 0 (heads in axils of ± foliaceous bracts; if peduncles produced, not inflated, strongly bracteate). Calyculi 0. Involucres campanulate to urceolate, 8–12[–16] mm diam. (larger in fruit). Phyllaries 24–30+ in 3+ series, lanceolate to lance-linear (± flat proximally), unequal to subequal, margins little, if at all, scarious, apices spine-tipped. Receptacles conic to hemispheric, smooth, glabrous, paleate; paleae winged (each palea enfolding, adnate to, and shed with cypsela). Florets 30–60+; corollas yellow. Cypselae (each shed with its enfolding palea) brownish, obcompressed, ± obovoid, not beaked, ribs 0, glabrous; pappi 0 or persistent (fragile), of 2–4[–5] whitish to stramineous, subequal, setiform-aristate scales plus 0–4 muticous to lanceolate scales in ± 1 series. x = 10.

Distribution

Europe, Middle East, n Africa, Atlantic Islands.

Discussion

Species 3 (2 in the flora).

Selected References

None.

Lower Taxa

Scolymus hispanicus, Scolymus maculatus

Key

1	Annuals; margins of stem wings and leaf blades usually white and thickened; pappi 0.	Scolymus maculatus
1	Biennials or perennials; margins of stem wings and leaf blades little, if at all, white or thickened; pappi of 2–4 setiform-aristate scales plus 0–4 muticous to lanceolate scales.	Scolymus hispanicus