Difference between revisions of "Selaginella ×neomexicana"

Maxon

Smithsonian Misc. Collect. 72: 2. 1920.

Treatment appears in FNA Volume 2.
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--><span class="statement" id="st-d0_s0" data-properties=""><b>Plants </b>on rock, forming clumps.</span> <span class="statement" id="st-d0_s1" data-properties="stem architecture or shape;stem location;stem location;stem shape"><b>Stems </b>radially symmetric, underground (rhizomatous) and aerial, not readily fragmenting, irregularly forked;</span> <span class="statement" id="st-d0_s2" data-properties="stem architecture;stem location;stem architecture or shape;branch count;tip course">rhizomatous and aerial stems often with 1 branch arrested, budlike, tips straight;</span> <span class="statement" id="st-d0_s3" data-properties="stem architecture;branch prominence;branch life cycle;branch architecture or shape">rhizomatous stems sometimes difficult to distinguish, without obvious living budlike branches;</span> <span class="statement" id="st-d0_s4" data-properties="stem location;stem orientation;stem orientation;stem orientation;branch architecture or shape">aerial stems erect to ascending, budlike branches mostly restricted to stem base (more conspicuous in ascending stems).</span> <span class="statement" id="st-d0_s5" data-properties="rhizophore position;rhizophore diam"><b>Rhizophores </b>borne on upperside of stems, restricted to lower 1/2 on erect stems or throughout stem length on ascending stems, 0.2–0.3 mm diam.</span> <span class="statement" id="st-d0_s6" data-properties="leaf growth form;leaf arrangement"><b>Leaves </b>dimorphic, not clearly ranked.</span> <span class="statement" id="st-d0_s7" data-properties="stem-leaf architecture;stem-leaf duration;stem-leaf duration;stem-leaf fusion;stem-leaf pubescence;stem location;stem location"><b>Rhizomatous </b>stem-leaves deciduous or persistent on base of emergent aerial stem, abruptly adnate, pubescent.</span> <span class="statement" id="st-d0_s8" data-properties="stem-leaf location;stem-leaf fixation or orientation;stem-leaf orientation;stem-leaf coloration;stem-leaf shape;stem-leaf length;stem-leaf width"><b>Aerial </b>stem-leaves appressed, ascending, green, linear-lanceolate, 1.9–2.7 X 0.36–0.46 mm;</span> <span class="statement" id="st-d0_s9" data-properties="abaxial ridge count">abaxial ridges present;</span> <span class="statement" id="st-d0_s10" data-properties="base fusion;base shape;base pubescence">base abruptly adnate, rounded, pubescent;</span> <span class="statement" id="st-d0_s11" data-properties="margin architecture or pubescence or shape;cilium coloration;cilium coloration;cilium coloration;cilium coloration;cilium coloration;cilium length or size;cilium orientation;cilium height or length or size;whole_organism some measurement">margins long-ciliate, cilia white, whitish to transparent or opaque, long and spreading at base, short to dentiform and ascending toward apex, 0.06–0.17 mm;</span> <span class="statement" id="st-d0_s12" data-properties="apex shape">apex keeled;</span> <span class="statement" id="st-d0_s13" data-properties="bristle coloration;bristle coloration;bristle coloration;bristle coloration;bristle coloration;bristle coloration;bristle coloration;bristle coloration;bristle coloration;bristle coloration;bristle pubescence;bristle some measurement">bristle whitish to white, greenish to yellowish opaque, slightly puberulent, 0.3–0.46 mm.</span> <span class="statement" id="st-d0_s14" data-properties="strobilus architecture or arrangement or growth form;strobilus atypical some measurement;strobilus some measurement"><b>Strobili </b>solitary, (0.5–) 1–3 cm;</span> <span class="statement" id="st-d0_s15" data-properties="sporophyll shape;sporophyll shape;sporophyll shape;abaxial ridge prominence;base pubescence;margin shape;apex shape;apex shape">sporophylls ovatelanceolate to lanceolate, abaxial ridges prominent, base glabrous, margins denticulate, apex keeled, short-bristled.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>on rock, forming clumps. <b>Stems</b> radially symmetric, underground (rhizomatous) and aerial, not readily fragmenting, irregularly forked; rhizomatous and aerial stems often with 1 branch arrested, budlike, tips straight; rhizomatous stems sometimes difficult to distinguish, without obvious living budlike branches; aerial stems erect to ascending, budlike branches mostly restricted to stem base (more conspicuous in ascending stems). <b>Rhizophores</b> borne on upperside of stems, restricted to lower 1/2 on erect stems or throughout stem length on ascending stems, 0.2–0.3 mm diam. <b>Leaves</b> dimorphic, not clearly ranked. <b>Rhizomatous</b> stem leaves deciduous or persistent on base of emergent aerial stem, abruptly adnate, pubescent. <b>Aerial</b> stem leaves appressed, ascending, green, linear-lanceolate, 1.9–2.7 × 0.36–0.46 mm; abaxial ridges present; base abruptly adnate, rounded, pubescent; margins long-ciliate, cilia white, whitish to transparent or opaque, long and spreading at base, short to dentiform and ascending toward apex, 0.06–0.17 mm; apex keeled; bristle whitish to white, greenish to yellowish opaque, slightly puberulent, 0.3–0.46 mm. <b>Strobili</b> solitary, (0.5–)1–3 cm; sporophylls ovate-lanceolate to lanceolate, abaxial ridges prominent, base glabrous, margins denticulate, apex keeled, short-bristled.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
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|elevation=1400–1700(–2000) m
 
|elevation=1400–1700(–2000) m
 
|distribution=Ariz.;N.Mex.;Tex.
 
|distribution=Ariz.;N.Mex.;Tex.
|discussion=<p>Selaginella × neomexicana is treated here as a hybrid, following R. M. Tryon (1955). Plants of this hybrid lack megaspores and megasporangia and have misshapen microsporangia. Several hypotheses for its origin have been advanced. It is clearly allied to S. rupincola, with which it shares white, long, spreading, marginal cilia on the leaves, hairs sometimes running along the ridges of the abaxial groove of the leaf, obscure rhizomatous underground stems, and buds mostly restricted to the base of aerial stems. Tryon (1955) suggested that the two presumed parents were S. rupincola and S. mutica, because S. × neomexicana has been found growing with S. mutica (usually var. limitanea). The usually strongly keeled apex in S. × neomexicana is a feature of S. mutica, and the range of S. × neomexicana is within the range of the two presumed parents. Selaginella underwoodii might conceivably be the second parent instead; its range overlaps the ranges of the putative hybrid and S. rupincola. It is possible that S. × neomexicana may represent an asexual race of S. rupincola. More detailed studies are necessary to determine the reproductive biology and cytology of this presumed hybrid and to assess its relationships.</p>
+
|discussion=<p><i>Selaginella</i> × <i>neomexicana</i> is treated here as a hybrid, following R. M. Tryon (1955). Plants of this hybrid lack megaspores and megasporangia and have misshapen microsporangia. Several hypotheses for its origin have been advanced. It is clearly allied to <i>S. rupincola</i>, with which it shares white, long, spreading, marginal cilia on the leaves, hairs sometimes running along the ridges of the abaxial groove of the leaf, obscure rhizomatous underground stems, and buds mostly restricted to the base of aerial stems. Tryon (1955) suggested that the two presumed parents were <i>S. rupincola</i> and <i>S. mutica</i>, because <i>S.</i> × <i>neomexicana</i> has been found growing with <i>S. mutica</i> (usually <i></i>var.<i> limitanea</i>). The usually strongly keeled apex in <i>S.</i> × <i>neomexicana</i> is a feature of <i>S. mutica</i>, and the range of <i>S.</i> × <i>neomexicana</i> is within the range of the two presumed parents. <i>Selaginella underwoodii</i> might conceivably be the second parent instead; its range overlaps the ranges of the putative hybrid and <i>S. rupincola</i>. It is possible that <i>S.</i> × <i>neomexicana</i> may represent an asexual race of <i>S. rupincola</i>. More detailed studies are necessary to determine the reproductive biology and cytology of this presumed hybrid and to assess its relationships.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Selaginella ×neomexicana
 
name=Selaginella ×neomexicana
|author=
 
 
|authority=Maxon
 
|authority=Maxon
 
|rank=species
 
|rank=species
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|publication year=1920
 
|publication year=1920
 
|special status=
 
|special status=
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_65.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_65.xml
 
|genus=Selaginella
 
|genus=Selaginella
 
|subgenus=Selaginella subg. Tetragonostachys
 
|subgenus=Selaginella subg. Tetragonostachys
 
|species=Selaginella ×neomexicana
 
|species=Selaginella ×neomexicana
|abaxial ridge count=present
 
|abaxial ridge prominence=prominent
 
|apex shape=short-bristled;keeled;keeled
 
|base fusion=adnate
 
|base pubescence=glabrous;pubescent
 
|base shape=rounded
 
|branch architecture or shape=budlike;budlike
 
|branch count=1
 
|branch life cycle=living
 
|branch prominence=obvious
 
|bristle coloration=whitish;white greenish
 
|bristle pubescence=puberulent
 
|bristle some measurement=0.3mm;0.46mm
 
|cilium coloration=whitish;transparent or opaque
 
|cilium height or length or size=short
 
|cilium length or size=long
 
|cilium orientation=spreading
 
|leaf arrangement=ranked
 
|leaf growth form=dimorphic
 
|margin architecture or pubescence or shape=long-ciliate
 
|margin shape=denticulate
 
|rhizophore diam=1/2
 
|rhizophore position=lower
 
|sporophyll shape=ovatelanceolate;lanceolate
 
|stem architecture=rhizomatous;rhizomatous
 
|stem architecture or shape=budlike;symmetric
 
|stem location=aerial;emergent;aerial;aerial;aerial;underground
 
|stem orientation=erect;ascending
 
|stem shape=forked
 
|stem-leaf architecture=rhizomatous
 
|stem-leaf coloration=green
 
|stem-leaf duration=persistent;deciduous
 
|stem-leaf fixation or orientation=appressed
 
|stem-leaf fusion=adnate
 
|stem-leaf length=1.9mm;2.7mm
 
|stem-leaf location=aerial
 
|stem-leaf orientation=ascending
 
|stem-leaf pubescence=pubescent
 
|stem-leaf shape=linear-lanceolate
 
|stem-leaf width=0.36mm;0.46mm
 
|strobilus architecture or arrangement or growth form=solitary
 
|strobilus atypical some measurement=0.5cm;1cm
 
|strobilus some measurement=1cm;3cm
 
|tip course=straight
 
|whole_organism some measurement=0.06mm;0.17mm
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Selaginella subg. Tetragonostachys]]
 
-->[[Category:Treatment]][[Category:Selaginella subg. Tetragonostachys]]

Latest revision as of 20:24, 5 November 2020

Plants on rock, forming clumps. Stems radially symmetric, underground (rhizomatous) and aerial, not readily fragmenting, irregularly forked; rhizomatous and aerial stems often with 1 branch arrested, budlike, tips straight; rhizomatous stems sometimes difficult to distinguish, without obvious living budlike branches; aerial stems erect to ascending, budlike branches mostly restricted to stem base (more conspicuous in ascending stems). Rhizophores borne on upperside of stems, restricted to lower 1/2 on erect stems or throughout stem length on ascending stems, 0.2–0.3 mm diam. Leaves dimorphic, not clearly ranked. Rhizomatous stem leaves deciduous or persistent on base of emergent aerial stem, abruptly adnate, pubescent. Aerial stem leaves appressed, ascending, green, linear-lanceolate, 1.9–2.7 × 0.36–0.46 mm; abaxial ridges present; base abruptly adnate, rounded, pubescent; margins long-ciliate, cilia white, whitish to transparent or opaque, long and spreading at base, short to dentiform and ascending toward apex, 0.06–0.17 mm; apex keeled; bristle whitish to white, greenish to yellowish opaque, slightly puberulent, 0.3–0.46 mm. Strobili solitary, (0.5–)1–3 cm; sporophylls ovate-lanceolate to lanceolate, abaxial ridges prominent, base glabrous, margins denticulate, apex keeled, short-bristled.


Habitat: On canyon rock
Elevation: 1400–1700(–2000) m

Discussion

Selaginella × neomexicana is treated here as a hybrid, following R. M. Tryon (1955). Plants of this hybrid lack megaspores and megasporangia and have misshapen microsporangia. Several hypotheses for its origin have been advanced. It is clearly allied to S. rupincola, with which it shares white, long, spreading, marginal cilia on the leaves, hairs sometimes running along the ridges of the abaxial groove of the leaf, obscure rhizomatous underground stems, and buds mostly restricted to the base of aerial stems. Tryon (1955) suggested that the two presumed parents were S. rupincola and S. mutica, because S. × neomexicana has been found growing with S. mutica (usually var. limitanea). The usually strongly keeled apex in S. × neomexicana is a feature of S. mutica, and the range of S. × neomexicana is within the range of the two presumed parents. Selaginella underwoodii might conceivably be the second parent instead; its range overlaps the ranges of the putative hybrid and S. rupincola. It is possible that S. × neomexicana may represent an asexual race of S. rupincola. More detailed studies are necessary to determine the reproductive biology and cytology of this presumed hybrid and to assess its relationships.

Selected References

None.

Lower Taxa

None.