Difference between revisions of "Coryphantha chlorantha"

(Engelmann) Britton & Rose

Cact. 4: 43. 1923.

Endemic
Basionym: Mammillaria chlorantha Engelmann in J. T. Rothrock, Rep. U.S. Geogr. Surv., Wheeler, 127. 1878
Synonyms: Coryphantha vivipara var. buoflama P. C. Fischer Coryphantha vivipara var. deserti (Engelmann) W. T. Marshall Escobaria deserti (Nuttall) Buxbaum Escobaria vivipara var. deserti (Engelmann) D. R. Hunt
Treatment appears in FNA Volume 4. Treatment on page 234. Mentioned on page 223, 235.
FNA>Volume Importer
 
imported>Volume Importer
 
(7 intermediate revisions by 2 users not shown)
Line 7: Line 7:
 
|year=1923
 
|year=1923
 
}}
 
}}
|basionyms={{Treatment/ID/Synonym
+
|special_status={{Treatment/ID/Special_status
 +
|code=E
 +
|label=Endemic
 +
}}
 +
|basionyms={{Treatment/ID/Basionym
 
|name=Mammillaria chlorantha
 
|name=Mammillaria chlorantha
 
|authority=Engelmann
 
|authority=Engelmann
 +
|rank=species
 +
|publication_title=in J. T. Rothrock, Rep. U.S. Geogr. Surv., Wheeler,
 +
|publication_place=127. 1878
 
}}
 
}}
 
|synonyms={{Treatment/ID/Synonym
 
|synonyms={{Treatment/ID/Synonym
 
|name=Coryphantha vivipara var. buoflama
 
|name=Coryphantha vivipara var. buoflama
 
|authority=P. C. Fischer
 
|authority=P. C. Fischer
}}{{Treatment/ID/Synonym
+
|rank=variety
 +
}} {{Treatment/ID/Synonym
 
|name=Coryphantha vivipara var. deserti
 
|name=Coryphantha vivipara var. deserti
 
|authority=(Engelmann) W. T. Marshall
 
|authority=(Engelmann) W. T. Marshall
}}{{Treatment/ID/Synonym
+
|rank=variety
 +
}} {{Treatment/ID/Synonym
 
|name=Escobaria deserti
 
|name=Escobaria deserti
 
|authority=(Nuttall) Buxbaum
 
|authority=(Nuttall) Buxbaum
}}{{Treatment/ID/Synonym
+
|rank=species
 +
}} {{Treatment/ID/Synonym
 
|name=Escobaria vivipara var. deserti
 
|name=Escobaria vivipara var. deserti
 
|authority=(Engelmann) D. R. Hunt
 
|authority=(Engelmann) D. R. Hunt
 +
|rank=variety
 
}}
 
}}
 
|hierarchy=Cactaceae;Cactaceae subfam. Cactoideae;Coryphantha;Coryphantha chlorantha
 
|hierarchy=Cactaceae;Cactaceae subfam. Cactoideae;Coryphantha;Coryphantha chlorantha
Line 31: Line 42:
 
}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="plant architecture;plant prominence;plant fragility;plant orientation;ground position relational"><b>Plants </b>usually unbranched (rarely to 12 branches), usually most protruding above ground level, stiff and strictly erect, densely covered with spines.</span> <span class="statement" id="st-d0_s1" data-properties="root density;taproot height or length or size;largest root count"><b>Roots </b>diffuse or short taproots, largest roots basally less than 1/5 of stem diam.</span> <span class="statement" id="st-d0_s2" data-properties="stem shape;stem shape;stem shape;stem shape;stem length;stem width"><b>Stems </b>spheric to short cylindric, 3.5–15 × 3.5–9.5 cm;</span> <span class="statement" id="st-d0_s3" data-properties="tubercle length;tubercle width;tubercle fragility">tubercles 9–14 × 4–7 mm, stiff;</span> <span class="statement" id="st-d0_s4" data-properties="areolar gland count">areolar glands absent;</span> <span class="statement" id="st-d0_s5" data-properties="parenchyma coating">parenchyma not mucilaginous;</span> <span class="statement" id="st-d0_s6" data-properties="druse size;druse some measurement;druse shape;cortex density">druses in pith and cortex sporadic, some giant, to 1 mm, lenticular;</span> <span class="statement" id="st-d0_s7" data-properties="pith count">pith 1/3–1/2 of lesser stem diam.;</span> <span class="statement" id="st-d0_s8" data-properties="medullary system architecture;medullary system prominence;medullary system density;medullary system count;plant life cycle">medullary vascular system conspicuous, diffuse, present even in immature plants.</span> <span class="statement" id="st-d0_s9" data-properties="spine count;spine coloration;spine coloration;spine coloration;spine coloration;spine coloration;spine reflectance;spine condition;spine coloration;tip coloration;tip count"><b>Spines </b>18–41 per areole, lightly pigmented, white to pale pinkish gray (varying geographically), brightest when fresh, weathering to gray, dark tips present on all or all but smallest spines;</span> <span class="statement" id="st-d0_s10" data-properties="spine arrangement;spine atypical count;spine count;spine atypical length;spine length;spine width">radial spines (12–) 15–33 per areole, 7.5–16.5 (–19) × 0.1–0.4 mm;</span> <span class="statement" id="st-d0_s11" data-properties="subcentral spine count;subcentral spine count">subcentral spines 1–7 often present adaxially;</span> <span class="statement" id="st-d0_s12" data-properties="outer central spine count">outer central spines 1–11 per areole;</span> <span class="statement" id="st-d0_s13" data-properties="inner central spine atypical count;inner central spine atypical count;inner central spine count;inner central spine orientation;inner central spine orientation;s-foot arrangement;largest spine life cycle;largest spine course;largest spine length;largest spine atypical width;largest spine width">inner central spines (0–) 1 (–2) per areole, porrect or ascending (other central spines usually not or weakly appressed), in bird’s-foot arrangement or radiating evenly in adults, straight, largest spines 13.5–20.5 × 0.3–0.6 (–0.8) mm.</span> <span class="statement" id="st-d0_s14" data-properties="flower length;flower width;flower position"><b>Flowers </b>nearly apical, 18–44 × 8–37 mm;</span> <span class="statement" id="st-d0_s15" data-properties="outer tepal shape">outer tepals densely fringed;</span> <span class="statement" id="st-d0_s16" data-properties="inner tepal count;inner tepal orientation;inner tepal orientation;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal coloration;inner tepal length;inner tepal width;midstripe prominence;midstripe coloration;mirroring coloration;mirroring coloration;mirroring coloration;mirroring coloration;mirroring coloration;mirroring reflectance;anther coloration;anther coloration;anther coloration;anther coloration;anther coloration;anther reflectance">inner tepals 13–34 per flower, ascending or recurved only at tips, pale dull creamy yellow or pale greenish tan to pale yellow-orange or pale apricot, usually with well-defined darker midstripes of maroon, magenta, pink, brownish orange, or yellow-green, proximally reflective, mirroring anthers, 11–25 × 1.8–2.8 mm;</span> <span class="statement" id="st-d0_s17" data-properties="outer filament coloration;outer filament coloration;outer filament coloration;outer filament coloration;outer filament coloration;outer filament position or shape;outer filament count">outer filaments white, pale-pink, or white with pinkish distal 1/2, not contrasting with inner tepals;</span> <span class="statement" id="st-d0_s18" data-properties="anther coloration">anthers bright orange-yellow;</span> <span class="statement" id="st-d0_s19" data-properties="stigma lobe count;stigma lobe orientation;stigma lobe orientation;stigma lobe coloration;stigma lobe coloration;stigma lobe coloration;part architecture;part coloration;part some measurement">stigma lobes 4–11, erect or ascending, white to greenish white, nonstigmatic parts slightly pink, 2.5–4.5 mm.</span> <span class="statement" id="st-d0_s20" data-properties="fruit coloration;part prominence;part reflectance;part coloration;part shape;part shape;part shape;part length;part width;part texture"><b>Fruits </b>pale green, exposed parts sometimes dull rose-pink, ellipsoid to ovoid, 12–31 × 5–11 mm, succulent;</span> <span class="statement" id="st-d0_s21" data-properties="fruit duration">floral remnant on fruit strongly persistent.</span> <span class="statement" id="st-d0_s22" data-properties=""><b>Seeds </b>reddish-brown, ± obovoid or comma-shaped, 1.4–1.8 mm, pitted.</span> <span class="statement" id="st-d0_s23" data-properties="seed coloration;seed shape;seed shape;seed some measurement;seed relief;2n chromosome count">2n = 22.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Plants </b>usually unbranched (rarely to 12 branches), usually most protruding above ground level, stiff and strictly erect, densely covered with spines. <b>Roots</b> diffuse or short taproots, largest roots basally less than 1/5 of stem diam. <b>Stems</b> spheric to short cylindric, 3.5–15 × 3.5–9.5 cm; tubercles 9–14 × 4–7 mm, stiff; areolar glands absent; parenchyma not mucilaginous; druses in pith and cortex sporadic, some giant, to 1 mm, lenticular; pith 1/3–1/2 of lesser stem diam.; medullary vascular system conspicuous, diffuse, present even in immature plants. <b>Spines</b> 18–41 per areole, lightly pigmented, white to pale pinkish gray (varying geographically), brightest when fresh, weathering to gray, dark tips present on all or all but smallest spines; radial spines (12–)15–33 per areole, 7.5–16.5(–19) × 0.1–0.4 mm; subcentral spines 1–7 often present adaxially; outer central spines 1–11 per areole; inner central spines (0–)1(–2) per areole, porrect or ascending (other central spines usually not or weakly appressed), in bird’s-foot arrangement or radiating evenly in adults, straight, largest spines 13.5–20.5 × 0.3–0.6(–0.8) mm. <b>Flowers</b> nearly apical, 18–44 × 8–37 mm; outer tepals densely fringed; inner tepals 13–34 per flower, ascending or recurved only at tips, pale dull creamy yellow or pale greenish tan to pale yellow-orange or pale apricot, usually with well-defined darker midstripes of maroon, magenta, pink, brownish orange, or yellow-green, proximally reflective, mirroring anthers, 11–25 × 1.8–2.8 mm; outer filaments white, pale pink, or white with pinkish distal 1/2, not contrasting with inner tepals; anthers bright orange-yellow; stigma lobes 4–11, erect or ascending, white to greenish white, nonstigmatic parts slightly pink, 2.5–4.5 mm. <b>Fruits</b> pale green, exposed parts sometimes dull rose-pink, ellipsoid to ovoid, 12–31 × 5–11 mm, succulent; floral remnant on fruit strongly persistent. <b>Seeds</b> reddish brown, ± obovoid or comma-shaped, 1.4–1.8 mm, pitted. <b>2n</b> = 22.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
 +
|phenology=Flowering Apr–Jun(-Sep); fruiting time unknown.
 
|habitat=Desert calcicolous scrub, Larrea scrub, blackbrush scrub, pinyon-juniper woodlands, limestone or dolomite rocky slopes, gravelly bajadas (rarely granite or basalt)
 
|habitat=Desert calcicolous scrub, Larrea scrub, blackbrush scrub, pinyon-juniper woodlands, limestone or dolomite rocky slopes, gravelly bajadas (rarely granite or basalt)
 
|elevation=(300-)500-1700(-2400) m
 
|elevation=(300-)500-1700(-2400) m
 
|distribution=Ariz.;Calif.;Nev.;Utah.
 
|distribution=Ariz.;Calif.;Nev.;Utah.
|discussion=<p>Coryphantha chlorantha is vegetatively identical to C. vivipara in the broad sense; without fresh flowers or reliable flower data, only an expert can distinguish C. chlorantha from C. vivipara, with which it sometimes grows sympatrically. Populations of C. chlorantha can be distinguished by their small, less-widely opening flowers, which are usually yellowish or pale pink-and-cream-striped, in contrast with the large, widely opening, purple flowers of C. vivipara.</p><!--
+
|discussion=<p><i>Coryphantha chlorantha</i> is vegetatively identical to <i>C. vivipara</i> in the broad sense; without fresh flowers or reliable flower data, only an expert can distinguish <i>C. chlorantha</i> from <i>C. vivipara</i>, with which it sometimes grows sympatrically. Populations of <i>C. chlorantha</i> can be distinguished by their small, less-widely opening flowers, which are usually yellowish or pale pink-and-cream-striped, in contrast with the large, widely opening, purple flowers of <i>C. vivipara</i>.</p><!--
--><p>Coryphantha chlorantha is more easily distinguished from C. alversonii, with which it is allopatric. Flowers of both are intermediate between those of C. vivipara and C. sneedii. Like C. sneedii, but unlike C. alversonii, C. chlorantha is polymorphic with respect to flower color, at least in some parts of its range. The “green” morph, for which chlorantha was named, probably is in the minority. A history of misidentifications limits the reliability of published photographs and distribution maps. The Arizona report of C. alversonii (L. D. Benson 1969) was based upon misidentified C. chlorantha or C. vivipara var. rosea.</p>
+
--><p><i>Coryphantha chlorantha</i> is more easily distinguished from <i>C. alversonii</i>, with which it is allopatric. Flowers of both are intermediate between those of <i>C. vivipara</i> and <i>C. sneedii</i>. Like <i>C. sneedii</i>, but unlike <i>C. alversonii</i>, <i>C. chlorantha</i> is polymorphic with respect to flower color, at least in some parts of its range. The “green” morph, for which chlorantha was named, probably is in the minority. A history of misidentifications limits the reliability of published photographs and distribution maps. The Arizona report of <i>C. alversonii</i> (L. D. Benson 1969) was based upon misidentified <i>C. chlorantha</i> or <i>C. vivipara</i> <i></i>var.<i> rosea</i>.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
Line 47: Line 59:
 
-->{{#Taxon:
 
-->{{#Taxon:
 
name=Coryphantha chlorantha
 
name=Coryphantha chlorantha
|author=
 
 
|authority=(Engelmann) Britton & Rose
 
|authority=(Engelmann) Britton & Rose
 
|rank=species
 
|rank=species
Line 54: Line 65:
 
|basionyms=Mammillaria chlorantha
 
|basionyms=Mammillaria chlorantha
 
|family=Cactaceae
 
|family=Cactaceae
 +
|phenology=Flowering Apr–Jun(-Sep); fruiting time unknown.
 
|habitat=Desert calcicolous scrub, Larrea scrub, blackbrush scrub, pinyon-juniper woodlands, limestone or dolomite rocky slopes, gravelly bajadas (rarely granite or basalt)
 
|habitat=Desert calcicolous scrub, Larrea scrub, blackbrush scrub, pinyon-juniper woodlands, limestone or dolomite rocky slopes, gravelly bajadas (rarely granite or basalt)
 
|elevation=(300-)500-1700(-2400) m
 
|elevation=(300-)500-1700(-2400) m
Line 60: Line 72:
 
|publication title=Cact.
 
|publication title=Cact.
 
|publication year=1923
 
|publication year=1923
|special status=
+
|special status=Endemic
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V4/V4_428.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V4/V4_428.xml
 
|subfamily=Cactaceae subfam. Cactoideae
 
|subfamily=Cactaceae subfam. Cactoideae
 
|genus=Coryphantha
 
|genus=Coryphantha
 
|species=Coryphantha chlorantha
 
|species=Coryphantha chlorantha
|2n chromosome count=22
 
|anther coloration=bright orange-yellow;yellow-green;brownish orange;pink;magenta;maroon
 
|anther reflectance=reflective
 
|areolar gland count=absent
 
|cortex density=sporadic
 
|druse shape=lenticular
 
|druse size=giant
 
|druse some measurement=0mm;1mm
 
|flower length=18mm;44mm
 
|flower position=apical
 
|flower width=8mm;37mm
 
|fruit coloration=pale green
 
|fruit duration=persistent
 
|ground position relational=level
 
|inner central spine atypical count=1;2
 
|inner central spine count=1
 
|inner central spine orientation=ascending;porrect
 
|inner tepal coloration=pale greenish tan;pale yellow-orange or pale apricot
 
|inner tepal count=13;34
 
|inner tepal length=11mm;25mm
 
|inner tepal orientation=recurved;ascending
 
|inner tepal width=1.8mm;2.8mm
 
|largest root count=0;1/5
 
|largest spine atypical width=0.6mm;0.8mm
 
|largest spine course=straight
 
|largest spine length=13.5mm;20.5mm
 
|largest spine life cycle=adults
 
|largest spine width=0.3mm;0.6mm
 
|medullary system architecture=vascular
 
|medullary system count=present
 
|medullary system density=diffuse
 
|medullary system prominence=conspicuous
 
|midstripe coloration=darker
 
|midstripe prominence=well-defined
 
|mirroring coloration=yellow-green;brownish orange;pink;magenta;maroon
 
|mirroring reflectance=reflective
 
|outer central spine count=1;11
 
|outer filament coloration=white with pinkish;pale-pink;white with pinkish;pale-pink;white
 
|outer filament count=1/2
 
|outer filament position or shape=distal
 
|outer tepal shape=fringed
 
|parenchyma coating=not mucilaginous
 
|part architecture=nonstigmatic
 
|part coloration=rose-pink;pink
 
|part length=12mm;31mm
 
|part prominence=exposed
 
|part reflectance=dull
 
|part shape=ellipsoid;ovoid
 
|part some measurement=2.5mm;4.5mm
 
|part texture=succulent
 
|part width=5mm;11mm
 
|pith count=1/3;1/2
 
|plant architecture=unbranched
 
|plant fragility=stiff
 
|plant life cycle=immature
 
|plant orientation=erect
 
|plant prominence=protruding
 
|root density=diffuse
 
|s-foot arrangement=radiating
 
|seed coloration=reddish-brown
 
|seed relief=pitted
 
|seed shape=comma-shaped;obovoid
 
|seed some measurement=1.4mm;1.8mm
 
|spine arrangement=radial
 
|spine atypical count=12;15
 
|spine atypical length=16.5mm;19mm
 
|spine coloration=gray;gray;white;pale pinkish
 
|spine condition=weathering
 
|spine count=15;33
 
|spine length=7.5mm;16.5mm
 
|spine reflectance=brightest
 
|spine width=0.1mm;0.4mm
 
|stem length=3.5cm;15cm
 
|stem shape=spheric;short cylindric
 
|stem width=3.5cm;9.5cm
 
|stigma lobe coloration=white;greenish white
 
|stigma lobe count=4;11
 
|stigma lobe orientation=ascending;erect
 
|subcentral spine count=present;1;7
 
|taproot height or length or size=short
 
|tip coloration=dark
 
|tip count=present
 
|tubercle fragility=stiff
 
|tubercle length=9mm;14mm
 
|tubercle width=4mm;7mm
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Coryphantha]]
 
-->[[Category:Treatment]][[Category:Coryphantha]]

Latest revision as of 21:58, 5 November 2020

Plants usually unbranched (rarely to 12 branches), usually most protruding above ground level, stiff and strictly erect, densely covered with spines. Roots diffuse or short taproots, largest roots basally less than 1/5 of stem diam. Stems spheric to short cylindric, 3.5–15 × 3.5–9.5 cm; tubercles 9–14 × 4–7 mm, stiff; areolar glands absent; parenchyma not mucilaginous; druses in pith and cortex sporadic, some giant, to 1 mm, lenticular; pith 1/3–1/2 of lesser stem diam.; medullary vascular system conspicuous, diffuse, present even in immature plants. Spines 18–41 per areole, lightly pigmented, white to pale pinkish gray (varying geographically), brightest when fresh, weathering to gray, dark tips present on all or all but smallest spines; radial spines (12–)15–33 per areole, 7.5–16.5(–19) × 0.1–0.4 mm; subcentral spines 1–7 often present adaxially; outer central spines 1–11 per areole; inner central spines (0–)1(–2) per areole, porrect or ascending (other central spines usually not or weakly appressed), in bird’s-foot arrangement or radiating evenly in adults, straight, largest spines 13.5–20.5 × 0.3–0.6(–0.8) mm. Flowers nearly apical, 18–44 × 8–37 mm; outer tepals densely fringed; inner tepals 13–34 per flower, ascending or recurved only at tips, pale dull creamy yellow or pale greenish tan to pale yellow-orange or pale apricot, usually with well-defined darker midstripes of maroon, magenta, pink, brownish orange, or yellow-green, proximally reflective, mirroring anthers, 11–25 × 1.8–2.8 mm; outer filaments white, pale pink, or white with pinkish distal 1/2, not contrasting with inner tepals; anthers bright orange-yellow; stigma lobes 4–11, erect or ascending, white to greenish white, nonstigmatic parts slightly pink, 2.5–4.5 mm. Fruits pale green, exposed parts sometimes dull rose-pink, ellipsoid to ovoid, 12–31 × 5–11 mm, succulent; floral remnant on fruit strongly persistent. Seeds reddish brown, ± obovoid or comma-shaped, 1.4–1.8 mm, pitted. 2n = 22.


Phenology: Flowering Apr–Jun(-Sep); fruiting time unknown.
Habitat: Desert calcicolous scrub, Larrea scrub, blackbrush scrub, pinyon-juniper woodlands, limestone or dolomite rocky slopes, gravelly bajadas (rarely granite or basalt)
Elevation: (300-)500-1700(-2400) m

Distribution

V4 428-distribution-map.gif

Ariz., Calif., Nev., Utah.

Discussion

Coryphantha chlorantha is vegetatively identical to C. vivipara in the broad sense; without fresh flowers or reliable flower data, only an expert can distinguish C. chlorantha from C. vivipara, with which it sometimes grows sympatrically. Populations of C. chlorantha can be distinguished by their small, less-widely opening flowers, which are usually yellowish or pale pink-and-cream-striped, in contrast with the large, widely opening, purple flowers of C. vivipara.

Coryphantha chlorantha is more easily distinguished from C. alversonii, with which it is allopatric. Flowers of both are intermediate between those of C. vivipara and C. sneedii. Like C. sneedii, but unlike C. alversonii, C. chlorantha is polymorphic with respect to flower color, at least in some parts of its range. The “green” morph, for which chlorantha was named, probably is in the minority. A history of misidentifications limits the reliability of published photographs and distribution maps. The Arizona report of C. alversonii (L. D. Benson 1969) was based upon misidentified C. chlorantha or C. vivipara var. rosea.

Selected References

None.

Lower Taxa

None.
... more about "Coryphantha chlorantha"
Allan D. Zimmerman +  and Bruce D. Parfitt +
(Engelmann) Britton & Rose +
Mammillaria chlorantha +
Ariz. +, Calif. +, Nev. +  and Utah. +
(300-)500-1700(-2400) m +
Desert calcicolous scrub, Larrea scrub, blackbrush scrub, pinyon-juniper woodlands, limestone or dolomite rocky slopes, gravelly bajadas (rarely granite or basalt) +
Flowering Apr–Jun(-Sep) +  and fruiting time unknown. +
Coryphantha vivipara var. buoflama +, Coryphantha vivipara var. deserti +, Escobaria deserti +  and Escobaria vivipara var. deserti +
Coryphantha chlorantha +
Coryphantha +
species +