Cercocarpus betuloides

Shrubs or trees, 10–50(–80) dm, sparsely to moderately branched, young stems, leaves, pedicels, and hypanthia sericeous (hairs straight, antrorse, appressed, or ± ascending), or hirsute-pilose (hairs erect, straight), or ± villous (hairs wavy or wavy-crinkled), or hairs tightly coiled or 1/2 coiled, or 1/2 coiled at base and erect-ascending, wavy distally, 0.2–0.8(–1.3) mm. Stems: long-shoot internodes 3–20(–55) mm, ± sparsely hairy, glabrescent; short shoots 3–30(–55) × 1–2.5 mm. Leaves persistent or ± drought-deciduous; stipules (1.5–)3.5–6(–10) mm, 1.5–2.5 mm in s California; petiole 1.5–9(–16) mm; blade oblong-obovate, rhombic, suborbiculate to broadly ovate, rarely oblong-elliptic, (8–)12–50(–78) × (4–)8–35(–64) mm, membranous to coriaceous, base cuneate to rounded, rarely ± cordate, margins ± revolute or flat, dentate, serrate, or, in broader leaves, crenate in distal 1/5–3/4, apex rounded, sometimes acute, abaxial surface sericeous, pilose, villous, or with curled hairs on veins, areoles canescent or glabrous, adaxial sparsely hairy, sometimes glabrate. Flowers 1–18 per short shoot; hypanthial tubes sericeous, woolly, or villous or with coiled or spreading hairs; hypanthial cups 2–4 × 4–7 mm; sepals 5, broadly deltate, 1–2.5 mm, acute-acuminate; stamens 20–61, anthers 1–1.5 mm, hirsute. Achenes 7–11(–13.5) × 1.5–2.5(–2.9) mm; fruiting pedicels (1.5–)3–11(–21) mm; hypanthial tubes (5–)7–13(–17) mm; pedicel/tube ratio (15–)35–90(–126)%; fruit awns 5–8.5(–12.5) cm, proximal setae 2.7–5 mm.

Distribution

• Generated Map
• Legacy Map

Ariz., Calif., Oreg., nw Mexico.

Discussion

Varieties 3 (3 in the flora).

The western, spring-flowering Cercocarpus betuloides in the broad sense, centered in California, is well supported by molecular data. A total of five taxa have been recognized in recent floras; molecular data provide no support for these taxa, but instead support recognition of a single inclusive taxon.

Selected References

None.

Lower Taxa

Cercocarpus betuloides var. betuloides, Cercocarpus betuloides var. blancheae, Cercocarpus betuloides var. traskiae

Key

1	Leaves, young stems, and hypanthial tubes appressed-sericeous or hypanthial tubes also with erect-ascending hairs (hairs not coiled); flowers 1–3(–10) per short shoot; transmontane s, c California, c Arizona, and sw Oregon.	Cercocarpus betuloides var. betuloides
1	Leaves, young stems, and hypanthial tubes commonly with tightly coiled hairs (except in some insular plants where sericeous); flowers (1–)3–6(–18) per short shoot; cismontane s California	> 2
2	Leaf blades sparsely to moderately hairy, subcoriaceous to coriaceous, margins serrate or dentate or crenate in distal 1/5–3/4; coastal s California and adjacent islands.	Cercocarpus betuloides var. blancheae
2	Leaf blades abaxially densely woolly, strongly coriaceous, margins (often strongly revolute, usually obscuring teeth), crenate, dentate-serrate in distal 1/2–3/4; sw Santa Catalina Island.	Cercocarpus betuloides var. traskiae