Difference between revisions of "Notholaena standleyi"

Maxon

Amer. Fern J. 5: 1. 1915.

Illustrated
Synonyms: Cheilanthes standleyi (Maxon) Mickel Chrysochosma hookeri (Domin) Kümmerle Notholaena hookeri D. C. Eaton
Treatment appears in FNA Volume 2.
FNA>Volume Importer
 
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|place=5: 1. 1915
 
|place=5: 1. 1915
 
|year=1915
 
|year=1915
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}}
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|special_status={{Treatment/ID/Special_status
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|code=F
 +
|label=Illustrated
 
}}
 
}}
 
|basionyms=
 
|basionyms=
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|name=Cheilanthes standleyi
 
|name=Cheilanthes standleyi
 
|authority=(Maxon) Mickel
 
|authority=(Maxon) Mickel
}}{{Treatment/ID/Synonym
+
|rank=species
 +
}} {{Treatment/ID/Synonym
 
|name=Chrysochosma hookeri
 
|name=Chrysochosma hookeri
 
|authority=(Domin) Kümmerle
 
|authority=(Domin) Kümmerle
}}{{Treatment/ID/Synonym
+
|rank=species
 +
}} {{Treatment/ID/Synonym
 
|name=Notholaena hookeri
 
|name=Notholaena hookeri
 
|authority=D. C. Eaton
 
|authority=D. C. Eaton
 +
|rank=species
 
}}
 
}}
 
|hierarchy=Pteridaceae;Notholaena;Notholaena standleyi
 
|hierarchy=Pteridaceae;Notholaena;Notholaena standleyi
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}}<!--
 
}}<!--
  
--><span class="statement" id="st-d0_s0" data-properties="stem scale coloration;margin coloration;margin width;margin prominence;margin width;margin architecture or shape"><b>Stem </b>scales strongly bicolored, margins brown, broad and well defined, thin, entire.</span> <span class="statement" id="st-d0_s1" data-properties="leaf some measurement"><b>Leaves </b>5–30 cm.</span> <span class="statement" id="st-d0_s2" data-properties="petiole coloration;petiole variability;petiole length or size;petiole shape;petiole pubescence;scale count"><b>Petiole </b>brown, equal to or somewhat longer than blade, rounded adaxially, glabrous except for a few scales at base.</span> <span class="statement" id="st-d0_s3" data-properties="blade shape;blade shape;blade architecture or shape;blade l w ratio;farina prominence;farina coloration;scale count;scale pubescence"><b>Blade </b>broadly pentagonal, deeply pinnatifid but not fully pinnate above base (distal pinnae connected by narrow wing of green tissue), 1–2 times longer than wide, abaxially with conspicuous yellowish farina, scales absent, adaxially glabrous;</span> <span class="statement" id="st-d0_s4" data-properties="basal pinna size;proximal pinna arrangement;proximal pinna shape;pinnule orientation;pinnule size">basal pinnae much larger than adjacent pair, strongly inequilateral, proximal basiscopic pinnules greatly enlarged.</span> <span class="statement" id="st-d0_s5" data-properties="ultimate segment architecture;ultimate segment fusion"><b>Ultimate </b>segments sessile, broadly adnate to costae;</span> <span class="statement" id="st-d0_s6" data-properties="segment margin orientation">segment margins slightly recurved, rarely concealing sporangia.</span> <span class="statement" id="st-d0_s7" data-properties=""><b>Sporangia </b>containing 32 or 16 spores.</span> <span class="statement" id="st-d0_s8" data-properties="sporangium count;sporangium count;2n chromosome count">2n = 60.</span><!--
+
--><span class="statement" id="st-undefined" data-properties=""><b>Stem </b>scales strongly bicolored, margins brown, broad and well defined, thin, entire. <b>Leaves</b> 5–30 cm. <b>Petiole</b> brown, equal to or somewhat longer than blade, rounded adaxially, glabrous except for a few scales at base. <b>Blade</b> broadly pentagonal, deeply pinnatifid but not fully pinnate above base (distal pinnae connected by narrow wing of green tissue), 1–2 times longer than wide, abaxially with conspicuous yellowish farina, scales absent, adaxially glabrous; basal pinnae much larger than adjacent pair, strongly inequilateral, proximal basiscopic pinnules greatly enlarged. <b>Ultimate</b> segments sessile, broadly adnate to costae; segment margins slightly recurved, rarely concealing sporangia. <b>Sporangia</b> containing 32 or 16 spores. <b>2n</b> = 60.</span><!--
  
 
-->{{Treatment/Body
 
-->{{Treatment/Body
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|phenology=Sporulating late spring–fall.
 
|habitat=Rocky slopes and cliffs, on a variety of substrates including granite and limestone
 
|habitat=Rocky slopes and cliffs, on a variety of substrates including granite and limestone
 
|elevation=300–2100 m
 
|elevation=300–2100 m
 
|distribution=Ariz.;Colo.;N.Mex.;Okla.;Tex.;Mexico.
 
|distribution=Ariz.;Colo.;N.Mex.;Okla.;Tex.;Mexico.
|discussion=<p>The low numbers of spores per sporangium in Notholaena standleyi are apparently not associated with apogamy because all plants thus far analyzed are sexual diploids (M. D. Windham, unpublished data). D. S. Siegler and E. Wollenweber (1983) identified three chemotypes in this species correlated with substrate specificity and subtle variations in farina color. These three "races" occupy different portions of the geographic range, and further investigation may indicate that they deserve formal taxonomic recognition. A report of Notholaena standleyi from Nevada (D. B. Lellinger 1985) must be considered suspect because it is disjunct, and the closest populations in Arizona represent a different chemotype.</p>
+
|discussion=<p>The low numbers of spores per sporangium in <i>Notholaena standleyi</i> are apparently not associated with apogamy because all plants thus far analyzed are sexual diploids (M. D. Windham, unpublished data). D. S. Siegler and E. Wollenweber (1983) identified three chemotypes in this species correlated with substrate specificity and subtle variations in farina color. These three "races" occupy different portions of the geographic range, and further investigation may indicate that they deserve formal taxonomic recognition. A report of <i>Notholaena standleyi</i> from <i>Nevada</i> (D. B. Lellinger 1985) must be considered suspect because it is disjunct, and the closest populations in Arizona represent a different chemotype.</p>
 
|tables=
 
|tables=
 
|references=
 
|references=
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-->{{#Taxon:
 
-->{{#Taxon:
 
name=Notholaena standleyi
 
name=Notholaena standleyi
|author=
 
 
|authority=Maxon
 
|authority=Maxon
 
|rank=species
 
|rank=species
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|basionyms=
 
|basionyms=
 
|family=Pteridaceae
 
|family=Pteridaceae
 +
|phenology=Sporulating late spring–fall.
 
|habitat=Rocky slopes and cliffs, on a variety of substrates including granite and limestone
 
|habitat=Rocky slopes and cliffs, on a variety of substrates including granite and limestone
 
|elevation=300–2100 m
 
|elevation=300–2100 m
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|publication title=Amer. Fern J.
 
|publication title=Amer. Fern J.
 
|publication year=1915
 
|publication year=1915
|special status=
+
|special status=Illustrated
|source xml=https://jpend@bitbucket.org/aafc-mbb/fna-fine-grained-xml.git/src/287ef3db526bd807d435a3c7423ef2df1e951227/V2/V2_705.xml
+
|source xml=https://bitbucket.org/aafc-mbb/fna-data-curation/src/2e0870ddd59836b60bcf96646a41e87ea5a5943a/coarse_grained_fna_xml/V2/V2_705.xml
 
|genus=Notholaena
 
|genus=Notholaena
 
|species=Notholaena standleyi
 
|species=Notholaena standleyi
|2n chromosome count=60
 
|basal pinna size=much larger
 
|blade architecture or shape=pinnate
 
|blade l w ratio=1-2
 
|blade shape=pinnatifid;pentagonal
 
|farina coloration=yellowish
 
|farina prominence=conspicuous
 
|leaf some measurement=5cm;30cm
 
|margin architecture or shape=entire
 
|margin coloration=brown
 
|margin prominence=defined
 
|margin width=thin;broad
 
|petiole coloration=brown
 
|petiole length or size=somewhat longer
 
|petiole pubescence=glabrous
 
|petiole shape=rounded
 
|petiole variability=equal
 
|pinnule orientation=basiscopic
 
|pinnule size=enlarged
 
|proximal pinna arrangement=adjacent
 
|proximal pinna shape=inequilateral
 
|scale count=absent;few
 
|scale pubescence=glabrous
 
|segment margin orientation=recurved
 
|sporangium count=16;32
 
|stem scale coloration=bicolored
 
|ultimate segment architecture=sessile
 
|ultimate segment fusion=adnate
 
 
}}<!--
 
}}<!--
  
 
-->[[Category:Treatment]][[Category:Notholaena]]
 
-->[[Category:Treatment]][[Category:Notholaena]]

Latest revision as of 20:24, 5 November 2020

Stem scales strongly bicolored, margins brown, broad and well defined, thin, entire. Leaves 5–30 cm. Petiole brown, equal to or somewhat longer than blade, rounded adaxially, glabrous except for a few scales at base. Blade broadly pentagonal, deeply pinnatifid but not fully pinnate above base (distal pinnae connected by narrow wing of green tissue), 1–2 times longer than wide, abaxially with conspicuous yellowish farina, scales absent, adaxially glabrous; basal pinnae much larger than adjacent pair, strongly inequilateral, proximal basiscopic pinnules greatly enlarged. Ultimate segments sessile, broadly adnate to costae; segment margins slightly recurved, rarely concealing sporangia. Sporangia containing 32 or 16 spores. 2n = 60.


Phenology: Sporulating late spring–fall.
Habitat: Rocky slopes and cliffs, on a variety of substrates including granite and limestone
Elevation: 300–2100 m

Distribution

V2 705-distribution-map.gif

Ariz., Colo., N.Mex., Okla., Tex., Mexico.

Discussion

The low numbers of spores per sporangium in Notholaena standleyi are apparently not associated with apogamy because all plants thus far analyzed are sexual diploids (M. D. Windham, unpublished data). D. S. Siegler and E. Wollenweber (1983) identified three chemotypes in this species correlated with substrate specificity and subtle variations in farina color. These three "races" occupy different portions of the geographic range, and further investigation may indicate that they deserve formal taxonomic recognition. A report of Notholaena standleyi from Nevada (D. B. Lellinger 1985) must be considered suspect because it is disjunct, and the closest populations in Arizona represent a different chemotype.

Selected References

None.

Lower Taxa

None.
... more about "Notholaena standleyi"
Michael D. Windham +
Ariz. +, Colo. +, N.Mex. +, Okla. +, Tex. +  and Mexico. +
300–2100 m +
Rocky slopes and cliffs, on a variety of substrates including granite and limestone +
Sporulating late spring–fall. +
Amer. Fern J. +
Illustrated +
Cheilanthes standleyi +, Chrysochosma hookeri +  and Notholaena hookeri +
Notholaena standleyi +
Notholaena +
species +