Psilostrophe

Biennials, perennials, subshrubs, or shrubs (rarely flowering first year), 8–60+ cm. Stems erect to spreading, branched from bases or throughout (not scapiform). Leaves basal and cauline or all cauline; alternate; petiolate or sessile; blades spatulate to oblanceolate or linear, margins usually entire (sometimes toothed or lobed on larger rosette leaves), faces densely to sparsely arachno-villous or ± strigillose, often gland-dotted as well. Heads radiate, usually in compact, corymbiform arrays or glomerulate clusters (borne singly in P. cooperi). Involucres cylindric to campanulate or obconic, 2–7 mm diam. Phyllaries persistent, 5–12 in 1–2 series (erect in fruit, distinct, oblong to lanceolate, bases ± indurate). Receptacles flat or convex, smooth or ± pitted (without setiform enations, sometimes gland-dotted), epaleate. Ray florets 1–8, pistillate, fertile; corollas yellow to orange (marcescent, spreading or reflexed in fruit). Disc florets 5–25+, bisexual, fertile; corollas yellow to orange, tubes shorter than narrowly cylindric throats, lobes 5, deltate (equal, papillate abaxially). Cypselae cylindric to clavate or obpyramidal, sometimes weakly obcompressed (ray), all striate-ribbed, usually glabrous, sometimes gland-dotted (villous in P. gnaphalodes, sometimes hirtellous in P. tagetina); pappi of 4–8 ± oblong or elliptic to lanceolate or lance-subulate, entire (lacerate in P. gnaphalodes) scales (without prominent midribs). x = 16.

Distribution

United States, Mexico.

Discussion

Species 7 (6 in the flora).

My treatment of psilostrophes closely follows that by R. C. Brown (1978). Psilostrophe mexicana R. C. Brown is known from Chihuahua and Durango.

Selected References

Brown, R. C. 1978. Biosystematics of Psilostrophe (Compositae: Helenieae). II. Artificial hybridization and systematic treatment. Madroño 25: 187–201.

Turner, B. L., M. W. Turner, and J. C. Crutchfield. 1988. Populational analyses and new combinations in Psilostrophe tagetina and P. gnaphalodes (Asteraceae, Heliantheae). Phytologia 65: 231–240.

Lower Taxa

Psilostrophe bakeri, Psilostrophe cooperi, Psilostrophe gnaphalodes, Psilostrophe sparsiflora, Psilostrophe tagetina, Psilostrophe villosa

Key

1	Subshrubs or shrubs; stems pannose (white); heads borne singly; peduncles 35–60(–80+) mm	Psilostrophe cooperi
1	Biennials or perennials; stems arachno-villous (gray to gray-green) or strigillose (greenish); heads in corymbiform arrays; peduncles 0.5–25(–50) mm	> 2
2	Stems ± strigillose, (greenish, sometimes ± villous in proximal axils); ray laminaereflexed in fruit	Psilostrophe sparsiflora
2	Stems mostly arachno-villous (gray to gray-green); ray laminae spreading in fruit	> 3
3	Involucres 7–9(–12) mm; disc florets (10–)12–15(–20); pappus scales oblong to ovate, 1.5–2 mm	Psilostrophe bakeri
3	Involucres 4–6(–8) mm; disc florets 5–8(–12); pappus scales elliptic or lanceolate to linear-subulate, 2–3+ mm	> 4
4	Cypselae villous	Psilostrophe gnaphalodes
4	Cypselae usually glabrous, sometimes hirtellous and/or gland-dotted	> 5
5	Peduncles (3–)12–20(–40) mm; ray laminae (5–)7–14+ mm	Psilostrophe tagetina
5	Peduncles (0.5–)1–3(–5+) mm; ray laminae 3–4(–6) mm	Psilostrophe villosa